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Sparks, John S. (2004). "Molecular phylogeny and biogeography of the Malagasy and South Asian cichlids (Teleostei: Perciformes: Cichlidae)". Molecular Phylogenetics and Evolution. 30 (3): 599–614. doi:10.1016/S1055-7903(03)00225-2. PMID15012941.