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Dutrillaux B.; Dutrillaux A.-M. (2013). "A South American Origin of the Genus Dynastes (Coleoptera: Scarabaeidae: Dynastinae) Demonstrated by Chromosomal Analyses". Cytogenetic and Genome Research. 147 (1): 37–42. doi:10.1159/000351210. PMID23735513. S2CID28561729.
Huang, J. (2016). "Parapatric genetic introgression and phenotypic assimilation: testing conditions for introgression between Hercules beetles (Dynastes, Dynastinae)". Molecular Ecology. 25 (21): 5513–5526. doi:10.1111/mec.13849. hdl:2027.42/134423. PMID27661063. S2CID22612044.
Dutrillaux B.; Dutrillaux A.-M. (2013). "A South American Origin of the Genus Dynastes (Coleoptera: Scarabaeidae: Dynastinae) Demonstrated by Chromosomal Analyses". Cytogenetic and Genome Research. 147 (1): 37–42. doi:10.1159/000351210. PMID23735513. S2CID28561729.