Humboldt penguin (English Wikipedia)

BirdLife International (2020). "Spheniscus humboldti". IUCN Red List of Threatened Species. 2020: e.T22697817A182714418. doi:10.2305/IUCN.UK.2020-3.RLTS.T22697817A182714418.en. Retrieved 13 November 2021.
Simeone, Alejandro; Schlatter, Roberto P. (1998). "Threats to a Mixed-Species Colony of Spheniscus Penguins in Southern Chile". Colonial Waterbirds. 21 (3): 418. doi:10.2307/1521654. JSTOR 1521654.
Ellenberg, Ursula; Mattern, Thomas; Seddon, Philip J.; Jorquera, Guillermo Luna (2006-11-01). "Physiological and reproductive consequences of human disturbance in Humboldt penguins: The need for species-specific visitor management". Biological Conservation. 133 (1): 95–106. Bibcode:2006BCons.133...95E. doi:10.1016/j.biocon.2006.05.019. ISSN 0006-3207.
Paredes, Rosana; Zavalaga, Carlos B.; Battistini, Gabriella; Majluf, Patricia; McGill, Patricia (2003). "Status of the Humboldt Penguin in Peru, 1999-2000". Waterbirds. 26 (2): 129. doi:10.1675/1524-4695(2003)026[0129:sothpi]2.0.co;2. ISSN 1524-4695. S2CID 85818196.
Paredes, Rosana; Zavalaga, Carlos B.; Boness, Daryl J. (2002-01-01). du Plessis, M. (ed.). "Patterns of Egg Laying and Breeding Success in Humboldt Penguins (Spheniscus Humboldti) at Punta San Juan, Peru". The Auk. 119 (1): 244–250. doi:10.1093/auk/119.1.244. ISSN 1938-4254.
Mattern, Thomas; Ellenberg, Ursula; Luna-Jorquera, Guillermo; Davis, Lloyd (2004-09-01). "Humboldt Penguin Census on Isla Chañaral, Chile: Recent Increase or Past Underestimate of Penguin Numbers?". Waterbirds. 27 (3): 268–276. doi:10.1675/1524-4695(2004)027[0368:HPCOIC]2.0.CO;2. S2CID 85668426.
Merritt, Kathleen; King, Nancy E. (1987). "Behavioral sex differences and activity patterns of captive humboldt penguins (Spheniscus humboldti)". Zoo Biology. 6 (2): 129–138. doi:10.1002/zoo.1430060204. ISSN 1098-2361.
Coker, Robert E. (1919). "Habits and economic relations of the guano birds of Peru" (PDF). Proceedings of the United States National Museum. 56 (2298): 449–511. doi:10.5479/si.00963801.56-2298.449. hdl:10088/15094. ISSN 0096-3801.
Culik, B.; Hennicke, J.; Martin, T. (2000-08-01). "Humboldt penguins outmanoeuvring El Nino". Journal of Experimental Biology. 203 (15): 2311–2322. doi:10.1242/jeb.203.15.2311. ISSN 0022-0949. PMID 10887069.
Hays, Coppelia (1986-01-01). "Effects of the 1982–1983 El Nino on Humboldt penguin colonies in Peru". Biological Conservation. 36 (2): 169–180. Bibcode:1986BCons..36..169H. doi:10.1016/0006-3207(86)90005-4. ISSN 0006-3207.
Wilson, Rory P.; Wilson, Marie-Pierre; Duffy, David Cameron; M, Braulio Araya; Klages, Norbert (1989-01-01). "Diving behaviour and prey of the Humboldt Penguin (Spheniscus humboldti)". Journal für Ornithologie. 130 (1): 75–79. Bibcode:1989JOrni.130...75W. doi:10.1007/BF01647164. ISSN 1439-0361. S2CID 38410399.
Herling, C.; Culik, B. M.; Hennicke, J. C. (May 2005). "Diet of the Humboldt penguin (Spheniscus humboldti) in northern and southern Chile". Marine Biology. 147 (1): 13–25. Bibcode:2005MarBi.147...13H. doi:10.1007/s00227-004-1547-8. ISSN 0025-3162. S2CID 55663752.
Martin, G.R.; Young, S.R. (1984-12-22). "The eye of the humboldt penguin, Spheniscus humboldti : visual fields and schematic optics". Proceedings of the Royal Society of London. Series B. Biological Sciences. 223 (1231): 197–222. Bibcode:1984RSPSB.223..197M. doi:10.1098/rspb.1984.0090. ISSN 0080-4649. PMID 6151660. S2CID 24295277.
Taylor, Sabrina S; Leonard, Marty L; Boness, Daryl J; Majluf, Patricia (2002-04-01). "Foraging by Humboldt penguins ( Spheniscus humboldti ) during the chick-rearing period: general patterns, sex differences, and recommendations to reduce incidental catches in fishing nets". Canadian Journal of Zoology. 80 (4): 700–707. doi:10.1139/z02-046. ISSN 0008-4301.
Williams, Tony D. (2012-08-05), "Parental Care", Physiological Adaptations for Breeding in Birds, Princeton University Press, doi:10.23943/princeton/9780691139821.003.0006, ISBN 978-0-691-13982-1
Blay, Nicola; Côté, Isabelle M. (December 2001). "Optimal conditions for breeding of captive humboldt penguins (Spheniscus humboldti): A survey of British zoos". Zoo Biology. 20 (6): 545–555. doi:10.1002/zoo.10002. ISSN 0733-3188.
Paredes, Rosana; Zavalaga, Carlos.B (August 2001). "Nesting sites and nest types as important factors for the conservation of Humboldt penguins (Sphensicus humboldti)". Biological Conservation. 100 (2): 199–205. Bibcode:2001BCons.100..199P. doi:10.1016/s0006-3207(01)00023-4. ISSN 0006-3207.
Hays, Coppelia (1986-01-01). "Effects of the 1982–1983 El Nino on Humboldt penguin colonies in Peru". Biological Conservation. 36 (2): 169–180. Bibcode:1986BCons..36..169H. doi:10.1016/0006-3207(86)90005-4. ISSN 0006-3207.
Majluf, Patricia; Babcock, Elizabeth A.; Riveros, Juan Carlos; Schreiber, Milena Arias; Alderete, William (October 2002). "Catch and Bycatch of Sea Birds and Marine Mammals in the Small-Scale Fishery of Punta San Juan, Peru". Conservation Biology. 16 (5): 1333–1343. Bibcode:2002ConBi..16.1333M. doi:10.1046/j.1523-1739.2002.00564.x. ISSN 0888-8892. S2CID 86452046.
Simeone, Alejandro; Bernal, Mariano (2000). "Effects of Habitat Modification on Breeding Seabirds: A Case Study in Central Chile". Waterbirds: The International Journal of Waterbird Biology. 23 (3): 449–456. doi:10.2307/1522182. ISSN 1524-4695. JSTOR 1522182.
Simeone, Alejandro; Luna-Jorquera, Guillermo (October 2012). "Estimating rat predation on Humboldt Penguin colonies in north-central Chile". Journal of Ornithology. 153 (4): 1079–1085. doi:10.1007/s10336-012-0837-z. hdl:10533/127889. ISSN 2193-7192. S2CID 11351679.
Paredes, Rosana; Zavalaga, Carlos B.; Battistini, Gabriella; Majluf, Patricia; McGill, Patricia (2003). "Status of the Humboldt Penguin in Peru, 1999-2000". Waterbirds. 26 (2): 129. doi:10.1675/1524-4695(2003)026[0129:sothpi]2.0.co;2. ISSN 1524-4695. S2CID 85818196.
Paredes, Rosana; Zavalaga, Carlos B.; Battistini, Gabriella; Majluf, Patricia; McGill, Patricia (2003). "Status of the Humboldt Penguin in Peru, 1999-2000". Waterbirds. 26 (2): 129. doi:10.1675/1524-4695(2003)026[0129:sothpi]2.0.co;2. ISSN 1524-4695. S2CID 85818196.

dublinzoo.ie

"Humboldt penguin". dublinzoo.ie. Dublin Zoo. Retrieved October 16, 2017.

fotawildlife.ie

"Humboldt penguin - Fota Wildlife Park". fotawildlife.ie. Fota Wildlife Park. Retrieved October 16, 2017.

handle.net

hdl.handle.net

Coker, Robert E. (1919). "Habits and economic relations of the guano birds of Peru" (PDF). Proceedings of the United States National Museum. 56 (2298): 449–511. doi:10.5479/si.00963801.56-2298.449. hdl:10088/15094. ISSN 0096-3801.
Simeone, Alejandro; Luna-Jorquera, Guillermo (October 2012). "Estimating rat predation on Humboldt Penguin colonies in north-central Chile". Journal of Ornithology. 153 (4): 1079–1085. doi:10.1007/s10336-012-0837-z. hdl:10533/127889. ISSN 2193-7192. S2CID 11351679.

harvard.edu

ui.adsabs.harvard.edu

Ellenberg, Ursula; Mattern, Thomas; Seddon, Philip J.; Jorquera, Guillermo Luna (2006-11-01). "Physiological and reproductive consequences of human disturbance in Humboldt penguins: The need for species-specific visitor management". Biological Conservation. 133 (1): 95–106. Bibcode:2006BCons.133...95E. doi:10.1016/j.biocon.2006.05.019. ISSN 0006-3207.
Hays, Coppelia (1986-01-01). "Effects of the 1982–1983 El Nino on Humboldt penguin colonies in Peru". Biological Conservation. 36 (2): 169–180. Bibcode:1986BCons..36..169H. doi:10.1016/0006-3207(86)90005-4. ISSN 0006-3207.
Wilson, Rory P.; Wilson, Marie-Pierre; Duffy, David Cameron; M, Braulio Araya; Klages, Norbert (1989-01-01). "Diving behaviour and prey of the Humboldt Penguin (Spheniscus humboldti)". Journal für Ornithologie. 130 (1): 75–79. Bibcode:1989JOrni.130...75W. doi:10.1007/BF01647164. ISSN 1439-0361. S2CID 38410399.
Herling, C.; Culik, B. M.; Hennicke, J. C. (May 2005). "Diet of the Humboldt penguin (Spheniscus humboldti) in northern and southern Chile". Marine Biology. 147 (1): 13–25. Bibcode:2005MarBi.147...13H. doi:10.1007/s00227-004-1547-8. ISSN 0025-3162. S2CID 55663752.
Martin, G.R.; Young, S.R. (1984-12-22). "The eye of the humboldt penguin, Spheniscus humboldti : visual fields and schematic optics". Proceedings of the Royal Society of London. Series B. Biological Sciences. 223 (1231): 197–222. Bibcode:1984RSPSB.223..197M. doi:10.1098/rspb.1984.0090. ISSN 0080-4649. PMID 6151660. S2CID 24295277.
Paredes, Rosana; Zavalaga, Carlos.B (August 2001). "Nesting sites and nest types as important factors for the conservation of Humboldt penguins (Sphensicus humboldti)". Biological Conservation. 100 (2): 199–205. Bibcode:2001BCons.100..199P. doi:10.1016/s0006-3207(01)00023-4. ISSN 0006-3207.
Hays, Coppelia (1986-01-01). "Effects of the 1982–1983 El Nino on Humboldt penguin colonies in Peru". Biological Conservation. 36 (2): 169–180. Bibcode:1986BCons..36..169H. doi:10.1016/0006-3207(86)90005-4. ISSN 0006-3207.
Majluf, Patricia; Babcock, Elizabeth A.; Riveros, Juan Carlos; Schreiber, Milena Arias; Alderete, William (October 2002). "Catch and Bycatch of Sea Birds and Marine Mammals in the Small-Scale Fishery of Punta San Juan, Peru". Conservation Biology. 16 (5): 1333–1343. Bibcode:2002ConBi..16.1333M. doi:10.1046/j.1523-1739.2002.00564.x. ISSN 0888-8892. S2CID 86452046.

independent.co.uk

"Penguins force suspension of $2.5bn mining project". The Independent. 2017-10-13. Retrieved 2020-07-10.
Farrell, Jeff (October 13, 2017). "Penguins put $2.5 billion mining project in Chile on ice". The Independent. Retrieved October 16, 2017.

iucnredlist.org

BirdLife International (2020). "Spheniscus humboldti". IUCN Red List of Threatened Species. 2020: e.T22697817A182714418. doi:10.2305/IUCN.UK.2020-3.RLTS.T22697817A182714418.en. Retrieved 13 November 2021.
"The IUCN red list of threatened species retrieved April 2019". IUCN Red List of Threatened Species.
BirdLife International (2018-08-09). "IUCN Red List of Threatened Species: Humboldt Penguin". IUCN Red List of Threatened Species. Retrieved 2020-04-08.
International), BirdLife International (BirdLife (2018-08-09). "IUCN Red List of Threatened Species: Spheniscus humboldti". IUCN Red List of Threatened Species. Retrieved 2020-07-10.

jstor.org

Simeone, Alejandro; Schlatter, Roberto P. (1998). "Threats to a Mixed-Species Colony of Spheniscus Penguins in Southern Chile". Colonial Waterbirds. 21 (3): 418. doi:10.2307/1521654. JSTOR 1521654.
Simeone, Alejandro; Bernal, Mariano (2000). "Effects of Habitat Modification on Breeding Seabirds: A Case Study in Central Chile". Waterbirds: The International Journal of Waterbird Biology. 23 (3): 449–456. doi:10.2307/1522182. ISSN 1524-4695. JSTOR 1522182.

luisenpark.de

"HUMBOLDT-PINGUIN". luisenpark.de. Luisenpark. Retrieved October 16, 2017.

marineornithology.org

"Changes in abundance and distribution of Humboldt Penguin. Viana et al, Marine Ornithology 42: 153–159 (2014)" (PDF).
Thumser, N.N.; Ficken, M.S. (1998). "Comparison of the vocal repertoires of captive Spheniscus penguins" (PDF). Marine Ornithology. 26: 41–48. Archived from the original (PDF) on 2021-04-10. Retrieved 2020-05-25.

milwaukeezoo.org

"Taylor Family Humboldt Penguin Exhibit". milwaukeezoo.org. Retrieved October 16, 2017.

mining-journal.com

"Andes Iron to fight for US$2.5B project". www.mining-journal.com. 2017-08-24. Retrieved 2020-07-10.

mining.com

"Andes Iron and Chilean ME enter conciliation process over $2.5-billion project". MINING.COM. 2018-02-12. Retrieved 2020-07-10.

mongabay.com

news.mongabay.com

"Chilean authorities eye controversial Cruz Grande port project". Mongabay Environmental News. 2020-03-25. Retrieved 2020-07-10.

newspapers.com

"Sea World's Split Personality (penguins, 1991)". Arizona Daily Star. 1991-06-23. p. 67. Retrieved 2020-07-02.
"Bronx Man Finds Penguin on Doorstep (1953)". The Troy Record. 1953-06-11. p. 31. Retrieved 2020-05-09.

nih.gov

pubmed.ncbi.nlm.nih.gov

Culik, B.; Hennicke, J.; Martin, T. (2000-08-01). "Humboldt penguins outmanoeuvring El Nino". Journal of Experimental Biology. 203 (15): 2311–2322. doi:10.1242/jeb.203.15.2311. ISSN 0022-0949. PMID 10887069.
Martin, G.R.; Young, S.R. (1984-12-22). "The eye of the humboldt penguin, Spheniscus humboldti : visual fields and schematic optics". Proceedings of the Royal Society of London. Series B. Biological Sciences. 223 (1231): 197–222. Bibcode:1984RSPSB.223..197M. doi:10.1098/rspb.1984.0090. ISSN 0080-4649. PMID 6151660. S2CID 24295277.

oup.com

academic.oup.com

Paredes, Rosana; Zavalaga, Carlos B.; Boness, Daryl J. (2002-01-01). du Plessis, M. (ed.). "Patterns of Egg Laying and Breeding Success in Humboldt Penguins (Spheniscus Humboldti) at Punta San Juan, Peru". The Auk. 119 (1): 244–250. doi:10.1093/auk/119.1.244. ISSN 1938-4254.

philadelphiazoo.org

"Humboldt penguin". Humboldt penguin - Philadelphia Zoo. Archived from the original on February 19, 2012. Retrieved October 16, 2017.

researchgate.net

Hiriart-Bertrand, L.; Simeone, A.; Reyes-Arriagada, R.; Riquelme, V.; Pütz, K.; Lüthi, B. (2010). "Description of a mixed-species colony of Humboldt (Spheniscus humboldti) and Magallanic Penguin (S. magellanicus) at Metalqui Island, Chiloé, southern Chile". Boletín Chileno de Ornitología. 16 (1): 42–47.
Zavalaga, C.B.; Paredes, R. (1997). "Sex determination of adult Humboldt penguins using morphometric characters". Journal of Field Ornithology. 68 (1): 102–112.
Mattern, Thomas; Ellenberg, Ursula; Luna-Jorquera, Guillermo; Davis, Lloyd (2004-09-01). "Humboldt Penguin Census on Isla Chañaral, Chile: Recent Increase or Past Underestimate of Penguin Numbers?". Waterbirds. 27 (3): 268–276. doi:10.1675/1524-4695(2004)027[0368:HPCOIC]2.0.CO;2. S2CID 85668426.
Valdés-Velasquez, Armando; de la Puente, Santiago; busalleu, alonso; cardeña, marco; Majluf, Patricia; Simeone, Alejandro (2013-01-01), "Humboldt Penguin (Spheniscus humboldti)", Penguins: Natural History and Conservation, p. 20, ISBN 978-0-295-99284-6, retrieved 2020-04-16

reuters.com

Fugitive penguin 337 spotted alive in Tokyo Bay, Reuters, 17 May 2012

sciencedirect.com

Ellenberg, Ursula; Mattern, Thomas; Seddon, Philip J.; Jorquera, Guillermo Luna (2006-11-01). "Physiological and reproductive consequences of human disturbance in Humboldt penguins: The need for species-specific visitor management". Biological Conservation. 133 (1): 95–106. Bibcode:2006BCons.133...95E. doi:10.1016/j.biocon.2006.05.019. ISSN 0006-3207.

seaturtles.org

Five Penguins Win U.S. Endangered Species Act Protection Turtle Island Restoration Network

semanticscholar.org

api.semanticscholar.org

Paredes, Rosana; Zavalaga, Carlos B.; Battistini, Gabriella; Majluf, Patricia; McGill, Patricia (2003). "Status of the Humboldt Penguin in Peru, 1999-2000". Waterbirds. 26 (2): 129. doi:10.1675/1524-4695(2003)026[0129:sothpi]2.0.co;2. ISSN 1524-4695. S2CID 85818196.
Mattern, Thomas; Ellenberg, Ursula; Luna-Jorquera, Guillermo; Davis, Lloyd (2004-09-01). "Humboldt Penguin Census on Isla Chañaral, Chile: Recent Increase or Past Underestimate of Penguin Numbers?". Waterbirds. 27 (3): 268–276. doi:10.1675/1524-4695(2004)027[0368:HPCOIC]2.0.CO;2. S2CID 85668426.
Wilson, Rory P.; Wilson, Marie-Pierre; Duffy, David Cameron; M, Braulio Araya; Klages, Norbert (1989-01-01). "Diving behaviour and prey of the Humboldt Penguin (Spheniscus humboldti)". Journal für Ornithologie. 130 (1): 75–79. Bibcode:1989JOrni.130...75W. doi:10.1007/BF01647164. ISSN 1439-0361. S2CID 38410399.
Herling, C.; Culik, B. M.; Hennicke, J. C. (May 2005). "Diet of the Humboldt penguin (Spheniscus humboldti) in northern and southern Chile". Marine Biology. 147 (1): 13–25. Bibcode:2005MarBi.147...13H. doi:10.1007/s00227-004-1547-8. ISSN 0025-3162. S2CID 55663752.
Martin, G.R.; Young, S.R. (1984-12-22). "The eye of the humboldt penguin, Spheniscus humboldti : visual fields and schematic optics". Proceedings of the Royal Society of London. Series B. Biological Sciences. 223 (1231): 197–222. Bibcode:1984RSPSB.223..197M. doi:10.1098/rspb.1984.0090. ISSN 0080-4649. PMID 6151660. S2CID 24295277.
Majluf, Patricia; Babcock, Elizabeth A.; Riveros, Juan Carlos; Schreiber, Milena Arias; Alderete, William (October 2002). "Catch and Bycatch of Sea Birds and Marine Mammals in the Small-Scale Fishery of Punta San Juan, Peru". Conservation Biology. 16 (5): 1333–1343. Bibcode:2002ConBi..16.1333M. doi:10.1046/j.1523-1739.2002.00564.x. ISSN 0888-8892. S2CID 86452046.
Simeone, Alejandro; Luna-Jorquera, Guillermo (October 2012). "Estimating rat predation on Humboldt Penguin colonies in north-central Chile". Journal of Ornithology. 153 (4): 1079–1085. doi:10.1007/s10336-012-0837-z. hdl:10533/127889. ISSN 2193-7192. S2CID 11351679.
Paredes, Rosana; Zavalaga, Carlos B.; Battistini, Gabriella; Majluf, Patricia; McGill, Patricia (2003). "Status of the Humboldt Penguin in Peru, 1999-2000". Waterbirds. 26 (2): 129. doi:10.1675/1524-4695(2003)026[0129:sothpi]2.0.co;2. ISSN 1524-4695. S2CID 85818196.
Paredes, Rosana; Zavalaga, Carlos B.; Battistini, Gabriella; Majluf, Patricia; McGill, Patricia (2003). "Status of the Humboldt Penguin in Peru, 1999-2000". Waterbirds. 26 (2): 129. doi:10.1675/1524-4695(2003)026[0129:sothpi]2.0.co;2. ISSN 1524-4695. S2CID 85818196.

si.edu

repository.si.edu

Coker, Robert E. (1919). "Habits and economic relations of the guano birds of Peru" (PDF). Proceedings of the United States National Museum. 56 (2298): 449–511. doi:10.5479/si.00963801.56-2298.449. hdl:10088/15094. ISSN 0096-3801.

tobuzoo.com

ペンギン舎. tobuzoo.com (in Japanese). Archived from the original on September 14, 2017. Retrieved October 16, 2017.

waza.org

"Humboldt Penguin - Spheniscus humboldti: WAZA: World Association of Zoos and Aquariums". WAZA. Archived from the original on 13 April 2012. Retrieved 12 February 2012.

web.archive.org

C. Michael Hogan (2008) Magellanic Penguin, GlobalTwitcher.com, ed. N. Stromberg Archived 2012-06-07 at the Wayback Machine
"Humboldt penguin". Humboldt penguin - Philadelphia Zoo. Archived from the original on February 19, 2012. Retrieved October 16, 2017.
"Humboldt Penguin - Spheniscus humboldti: WAZA: World Association of Zoos and Aquariums". WAZA. Archived from the original on 13 April 2012. Retrieved 12 February 2012.
Thumser, N.N.; Ficken, M.S. (1998). "Comparison of the vocal repertoires of captive Spheniscus penguins" (PDF). Marine Ornithology. 26: 41–48. Archived from the original (PDF) on 2021-04-10. Retrieved 2020-05-25.
ペンギン舎. tobuzoo.com (in Japanese). Archived from the original on September 14, 2017. Retrieved October 16, 2017.
"Humboldt penguins". chesterzoo.org. Archived from the original on October 16, 2017. Retrieved October 16, 2017.

worldcat.org

search.worldcat.org

Ellenberg, Ursula; Mattern, Thomas; Seddon, Philip J.; Jorquera, Guillermo Luna (2006-11-01). "Physiological and reproductive consequences of human disturbance in Humboldt penguins: The need for species-specific visitor management". Biological Conservation. 133 (1): 95–106. Bibcode:2006BCons.133...95E. doi:10.1016/j.biocon.2006.05.019. ISSN 0006-3207.
Paredes, Rosana; Zavalaga, Carlos B.; Battistini, Gabriella; Majluf, Patricia; McGill, Patricia (2003). "Status of the Humboldt Penguin in Peru, 1999-2000". Waterbirds. 26 (2): 129. doi:10.1675/1524-4695(2003)026[0129:sothpi]2.0.co;2. ISSN 1524-4695. S2CID 85818196.
Paredes, Rosana; Zavalaga, Carlos B.; Boness, Daryl J. (2002-01-01). du Plessis, M. (ed.). "Patterns of Egg Laying and Breeding Success in Humboldt Penguins (Spheniscus Humboldti) at Punta San Juan, Peru". The Auk. 119 (1): 244–250. doi:10.1093/auk/119.1.244. ISSN 1938-4254.
Merritt, Kathleen; King, Nancy E. (1987). "Behavioral sex differences and activity patterns of captive humboldt penguins (Spheniscus humboldti)". Zoo Biology. 6 (2): 129–138. doi:10.1002/zoo.1430060204. ISSN 1098-2361.
Coker, Robert E. (1919). "Habits and economic relations of the guano birds of Peru" (PDF). Proceedings of the United States National Museum. 56 (2298): 449–511. doi:10.5479/si.00963801.56-2298.449. hdl:10088/15094. ISSN 0096-3801.
Culik, B.; Hennicke, J.; Martin, T. (2000-08-01). "Humboldt penguins outmanoeuvring El Nino". Journal of Experimental Biology. 203 (15): 2311–2322. doi:10.1242/jeb.203.15.2311. ISSN 0022-0949. PMID 10887069.
Hays, Coppelia (1986-01-01). "Effects of the 1982–1983 El Nino on Humboldt penguin colonies in Peru". Biological Conservation. 36 (2): 169–180. Bibcode:1986BCons..36..169H. doi:10.1016/0006-3207(86)90005-4. ISSN 0006-3207.
Wilson, Rory P.; Wilson, Marie-Pierre; Duffy, David Cameron; M, Braulio Araya; Klages, Norbert (1989-01-01). "Diving behaviour and prey of the Humboldt Penguin (Spheniscus humboldti)". Journal für Ornithologie. 130 (1): 75–79. Bibcode:1989JOrni.130...75W. doi:10.1007/BF01647164. ISSN 1439-0361. S2CID 38410399.
Herling, C.; Culik, B. M.; Hennicke, J. C. (May 2005). "Diet of the Humboldt penguin (Spheniscus humboldti) in northern and southern Chile". Marine Biology. 147 (1): 13–25. Bibcode:2005MarBi.147...13H. doi:10.1007/s00227-004-1547-8. ISSN 0025-3162. S2CID 55663752.
Martin, G.R.; Young, S.R. (1984-12-22). "The eye of the humboldt penguin, Spheniscus humboldti : visual fields and schematic optics". Proceedings of the Royal Society of London. Series B. Biological Sciences. 223 (1231): 197–222. Bibcode:1984RSPSB.223..197M. doi:10.1098/rspb.1984.0090. ISSN 0080-4649. PMID 6151660. S2CID 24295277.
Taylor, Sabrina S; Leonard, Marty L; Boness, Daryl J; Majluf, Patricia (2002-04-01). "Foraging by Humboldt penguins ( Spheniscus humboldti ) during the chick-rearing period: general patterns, sex differences, and recommendations to reduce incidental catches in fishing nets". Canadian Journal of Zoology. 80 (4): 700–707. doi:10.1139/z02-046. ISSN 0008-4301.
Blay, Nicola; Côté, Isabelle M. (December 2001). "Optimal conditions for breeding of captive humboldt penguins (Spheniscus humboldti): A survey of British zoos". Zoo Biology. 20 (6): 545–555. doi:10.1002/zoo.10002. ISSN 0733-3188.
Paredes, Rosana; Zavalaga, Carlos.B (August 2001). "Nesting sites and nest types as important factors for the conservation of Humboldt penguins (Sphensicus humboldti)". Biological Conservation. 100 (2): 199–205. Bibcode:2001BCons.100..199P. doi:10.1016/s0006-3207(01)00023-4. ISSN 0006-3207.
Hays, Coppelia (1986-01-01). "Effects of the 1982–1983 El Nino on Humboldt penguin colonies in Peru". Biological Conservation. 36 (2): 169–180. Bibcode:1986BCons..36..169H. doi:10.1016/0006-3207(86)90005-4. ISSN 0006-3207.
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