Humulus lupulus (English Wikipedia)

Conway, Sean; Snyder, Reid (2008). "Humulus lupulus - Hops" (PDF). College Seminar 235 Food for Thought: The Science, Culture, & Politics of Food. Hamilton College. Retrieved July 31, 2008.

hdl.handle.net (Global: 66^th place; English: 34^th place)

Zanoli, Paola; Zavatti, Manuela (2008). "Pharmacognostic and pharmacological profile of Humulus lupulus L". Journal of Ethnopharmacology. 116 (3): 383–396. doi:10.1016/j.jep.2008.01.011. hdl:11380/611737. PMID 18308492.

herbalgram.org (Global: low place; English: low place)

cms.herbalgram.org

"HerbalGram: Hops (Humulus lupulus): A Review of its Historic and Medicinal Uses". cms.herbalgram.org. Retrieved 2017-04-08.^{[permanent dead link]}

iucnredlist.org (Global: 39^th place; English: 71^st place)

Khela, S. (2012). "Humulus lupulus (Europe assessment)". IUCN Red List of Threatened Species. 2012 e.T202971A2758327. Retrieved 9 November 2025.

luontoportti.com (Global: 8,871^st place; English: low place)

"Hop, Humulus lupulus - Flowers". NatureGate. Retrieved 2017-04-13.

missouribotanicalgarden.org (Global: 5,451^st place; English: 3,624^th place)

"Humulus lupulus". Plant Finder. Missouri botanical Garden. Retrieved 2017-04-12.

nativeplanttrust.org (Global: low place; English: low place)

gobotany.nativeplanttrust.org

"Humulus lupulus (common hop, hops)". Go Botany. New England Wildflower Society. Retrieved 2017-04-13.

natureserve.org (Global: 2,387^th place; English: 1,112^th place)

explorer.natureserve.org

NatureServe. "Humulus lupulus". NatureServe Explorer. Arlington, Virginia. Retrieved 9 November 2025.

nih.gov (Global: 5^th place; English: 5^th place)

pubmed.ncbi.nlm.nih.gov

Zanoli, Paola; Zavatti, Manuela (2008). "Pharmacognostic and pharmacological profile of Humulus lupulus L". Journal of Ethnopharmacology. 116 (3): 383–396. doi:10.1016/j.jep.2008.01.011. hdl:11380/611737. PMID 18308492.
Eriksen, Renée L.; Padgitt-Cobb, Lillian K.; Townsend, M. Shaun; Henning, John A. (December 2021). "Gene expression for secondary metabolite biosynthesis in hop (Humulus lupulus L.) leaf lupulin glands exposed to heat and low-water stress". Scientific Reports. 11 (1): 5138. Bibcode:2021NatSR..11.5138E. doi:10.1038/s41598-021-84691-y. ISSN 2045-2322. PMC 7970847. PMID 33664420.
Goese, Markus; Kammhuber, Klaus; Bacher, Adelbert; Zenk, Meinhart H.; Eisenreich, Wolfgang (1999-07-15). "Biosynthesis of bitter acids in hops. A 13C-NMR and 2H-NMR study on the building blocks of humulone". European Journal of Biochemistry. 263 (2): 447–454. Bibcode:1999EJBio.263..447G. doi:10.1046/j.1432-1327.1999.00518.x. ISSN 0014-2956. PMID 10406953.
Natsume, S.; Takagi, H.; Shiraishi, A.; Murata, J.; Toyonaga, H.; Patzak, J.; Takagi, M.; Yaegashi, H.; Uemura, A.; Mitsuoka, C.; Yoshida, K. (2014-11-20). "The Draft Genome of Hop (Humulus lupulus), an Essence for Brewing". Plant and Cell Physiology. 56 (3): 428–441. doi:10.1093/pcp/pcu169. ISSN 0032-0781. PMID 25416290.
Padgitt-Cobb, Lillian K.; Kingan, Sarah B.; Wells, Jackson; Elser, Justin; Kronmiller, Brent; Moore, Daniel; Concepcion, Gregory; Peluso, Paul; Rank, David; Jaiswal, Pankaj; Henning, John; Hendrix, David A. (2021-02-18). "A draft phased assembly of the diploid Cascade hop ( Humulus lupulus ) genome". The Plant Genome. 14 (1) e20072. doi:10.1002/tpg2.20072. ISSN 1940-3372. PMID 33605092. S2CID 231962731.
Milligan SR, Kalita JC, Heyerick A, Rong H, De Cooman L, De Keukeleire D (June 1999). "Identification of a potent phytoestrogen in hops (Humulus lupulus L.) and beer". J Clin Endocrinol Metab. 84 (6): 2249–52. doi:10.1210/jcem.84.6.5887. PMID 10372741.
Benkherouf, Ali Y.; Soini, Sanna L.; Stompor, Monika; Uusi-Oukari, Mikko (February 2019). "Positive allosteric modulation of native and recombinant GABAA receptors by hops prenylflavonoids". European Journal of Pharmacology. 852: 34–41. doi:10.1016/j.ejphar.2019.02.034. ISSN 0014-2999. PMID 30797788. S2CID 73456325.
Rozalski M, Micota B, Sadowska B, Stochmal A, Jedrejek D, Wieckowska-Szakiel M, Rozalska B (2013). "Antiadherent and Antibiofilm Activity of Humulus lupulus L. Derived Products: New Pharmacological Properties". BioMed Research International. 2013: 1–7. doi:10.1155/2013/101089. PMC 3794639. PMID 24175280.
Franco, L.; Sánchez, C.; Bravo, R.; Rodriguez, A.; Barriga, C.; Juánez, Javier Cubero (2012-06-01). "The sedative effects of hops (Humulus lupulus), a component of beer, on the activity/rest rhythm". Acta Physiologica Hungarica. 99 (2): 133–139. doi:10.1556/APhysiol.99.2012.2.6. ISSN 0231-424X. PMID 22849837.
Divashuk, Mikhail G.; Alexandrov, Oleg S.; Razumova, Olga V.; Kirov, Ilya V.; Karlov, Gennady I. (2014-01-21). "Molecular Cytogenetic Characterization of the Dioecious Cannabis sativa with an XY Chromosome Sex Determination System". PLOS ONE. 9 (1) e85118. Bibcode:2014PLoSO...985118D. doi:10.1371/journal.pone.0085118. ISSN 1932-6203. PMC 3897423. PMID 24465491.
Murakami, A; Darby, P; Javornik, B; Pais, M S S; Seigner, E; Lutz, A; Svoboda, P (2006-05-10). "Molecular phylogeny of wild Hops, Humulus lupulus L." Heredity. 97 (1): 66–74. Bibcode:2006Hered..97...66M. doi:10.1038/sj.hdy.6800839. ISSN 0018-067X. PMID 16685279. S2CID 11920277.
Pisupati, Rahul; Vergara, Daniela; Kane, Nolan C. (2018-02-21). "Diversity and evolution of the repetitive genomic content in Cannabis sativa". BMC Genomics. 19 (1): 156. doi:10.1186/s12864-018-4494-3. ISSN 1471-2164. PMC 5822635. PMID 29466945.
Laverty, Kaitlin U.; Stout, Jake M.; Sullivan, Mitchell J.; Shah, Hardik; Gill, Navdeep; Holbrook, Larry; Deikus, Gintaras; Sebra, Robert; Hughes, Timothy R.; Page, Jonathan E.; van Bakel, Harm (2018-11-08). "A physical and genetic map of Cannabis sativa identifies extensive rearrangements at the THC/CBD acid synthase loci". Genome Research. 29 (1): 146–156. doi:10.1101/gr.242594.118. ISSN 1088-9051. PMC 6314170. PMID 30409771.
Marks, M. David; Tian, Li; Wenger, Jonathan P.; Omburo, Stephanie N.; Soto-Fuentes, Wilfredo; He, Ji; Gang, David R.; Weiblen, George D.; Dixon, Richard A. (2009-07-06). "Identification of candidate genes affecting Δ9-tetrahydrocannabinol biosynthesis in Cannabis sativa". Journal of Experimental Botany. 60 (13): 3715–3726. doi:10.1093/jxb/erp210. ISSN 1460-2431. PMC 2736886. PMID 19581347.
Langezaal CR, Chandra A, Scheffer JJ (1992). "Antimicrobial screening of essential oils and extracts of some Humulus lupulus L. cultivars". Pharm Weekbl Sci. 14 (6): 353–356. doi:10.1007/bf01970171. PMID 1475174. S2CID 12561634.
Buso, Piergiacomo; Manfredini, Stefano; Reza Ahmadi-Ashtiani, Hamid; Sciabica, Sabrina; Buzzi, Raissa; Vertuani, Silvia; Baldisserotto, Anna (26 February 2020). "Iranian Medicinal Plants: From Ethnomedicine to Actual Studies". Medicina. 56 (3): 18. doi:10.3390/medicina56030097. PMC 7143749. PMID 32110920.

ncbi.nlm.nih.gov

Eriksen, Renée L.; Padgitt-Cobb, Lillian K.; Townsend, M. Shaun; Henning, John A. (December 2021). "Gene expression for secondary metabolite biosynthesis in hop (Humulus lupulus L.) leaf lupulin glands exposed to heat and low-water stress". Scientific Reports. 11 (1): 5138. Bibcode:2021NatSR..11.5138E. doi:10.1038/s41598-021-84691-y. ISSN 2045-2322. PMC 7970847. PMID 33664420.
Rozalski M, Micota B, Sadowska B, Stochmal A, Jedrejek D, Wieckowska-Szakiel M, Rozalska B (2013). "Antiadherent and Antibiofilm Activity of Humulus lupulus L. Derived Products: New Pharmacological Properties". BioMed Research International. 2013: 1–7. doi:10.1155/2013/101089. PMC 3794639. PMID 24175280.
Divashuk, Mikhail G.; Alexandrov, Oleg S.; Razumova, Olga V.; Kirov, Ilya V.; Karlov, Gennady I. (2014-01-21). "Molecular Cytogenetic Characterization of the Dioecious Cannabis sativa with an XY Chromosome Sex Determination System". PLOS ONE. 9 (1) e85118. Bibcode:2014PLoSO...985118D. doi:10.1371/journal.pone.0085118. ISSN 1932-6203. PMC 3897423. PMID 24465491.
Pisupati, Rahul; Vergara, Daniela; Kane, Nolan C. (2018-02-21). "Diversity and evolution of the repetitive genomic content in Cannabis sativa". BMC Genomics. 19 (1): 156. doi:10.1186/s12864-018-4494-3. ISSN 1471-2164. PMC 5822635. PMID 29466945.
Laverty, Kaitlin U.; Stout, Jake M.; Sullivan, Mitchell J.; Shah, Hardik; Gill, Navdeep; Holbrook, Larry; Deikus, Gintaras; Sebra, Robert; Hughes, Timothy R.; Page, Jonathan E.; van Bakel, Harm (2018-11-08). "A physical and genetic map of Cannabis sativa identifies extensive rearrangements at the THC/CBD acid synthase loci". Genome Research. 29 (1): 146–156. doi:10.1101/gr.242594.118. ISSN 1088-9051. PMC 6314170. PMID 30409771.
Marks, M. David; Tian, Li; Wenger, Jonathan P.; Omburo, Stephanie N.; Soto-Fuentes, Wilfredo; He, Ji; Gang, David R.; Weiblen, George D.; Dixon, Richard A. (2009-07-06). "Identification of candidate genes affecting Δ9-tetrahydrocannabinol biosynthesis in Cannabis sativa". Journal of Experimental Botany. 60 (13): 3715–3726. doi:10.1093/jxb/erp210. ISSN 1460-2431. PMC 2736886. PMID 19581347.
Buso, Piergiacomo; Manfredini, Stefano; Reza Ahmadi-Ashtiani, Hamid; Sciabica, Sabrina; Buzzi, Raissa; Vertuani, Silvia; Baldisserotto, Anna (26 February 2020). "Iranian Medicinal Plants: From Ethnomedicine to Actual Studies". Medicina. 56 (3): 18. doi:10.3390/medicina56030097. PMC 7143749. PMID 32110920.

plantlife.org.uk (Global: low place; English: low place)

Plantlife website County Flowers page Archived 2015-04-30 at the Wayback Machine

purdue.edu (Global: 1,391^st place; English: 979^th place)

hort.purdue.edu

"Humulus lupulus". NewCROP, the New Crop Resource Online Program. Retrieved 2017-04-02.

researchgate.net (Global: 96^th place; English: 89^th place)

Behre, Karl-Ernst (1999). "The history of beer additives in Europe - a review". Vegetation History and Archaeobotany. 8 (1–2): 35–48. Bibcode:1999VegHA...8...35B. doi:10.1007/BF02042841. S2CID 129429552.

rhs.org.uk (Global: 1,321^st place; English: 666^th place)

"Humulus lupulus 'Aureus' AGM". RHS Plant Selector. Royal Horticultural Society. Retrieved 25 August 2020.
"AGM Plants - Ornamental" (PDF). Royal Horticultural Society. July 2017. p. 43. Retrieved 3 March 2018.

sciencedaily.com (Global: 1,390^th place; English: 1,286^th place)

"Advancing understanding of hop genome to aid brewers, medical researchers". ScienceDaily. Retrieved 2021-04-09.

semanticscholar.org (Global: 15^th place; English: 8^th place)

api.semanticscholar.org

Padgitt-Cobb, Lillian K.; Kingan, Sarah B.; Wells, Jackson; Elser, Justin; Kronmiller, Brent; Moore, Daniel; Concepcion, Gregory; Peluso, Paul; Rank, David; Jaiswal, Pankaj; Henning, John; Hendrix, David A. (2021-02-18). "A draft phased assembly of the diploid Cascade hop ( Humulus lupulus ) genome". The Plant Genome. 14 (1) e20072. doi:10.1002/tpg2.20072. ISSN 1940-3372. PMID 33605092. S2CID 231962731.
Benkherouf, Ali Y.; Soini, Sanna L.; Stompor, Monika; Uusi-Oukari, Mikko (February 2019). "Positive allosteric modulation of native and recombinant GABAA receptors by hops prenylflavonoids". European Journal of Pharmacology. 852: 34–41. doi:10.1016/j.ejphar.2019.02.034. ISSN 0014-2999. PMID 30797788. S2CID 73456325.
Murakami, A; Darby, P; Javornik, B; Pais, M S S; Seigner, E; Lutz, A; Svoboda, P (2006-05-10). "Molecular phylogeny of wild Hops, Humulus lupulus L." Heredity. 97 (1): 66–74. Bibcode:2006Hered..97...66M. doi:10.1038/sj.hdy.6800839. ISSN 0018-067X. PMID 16685279. S2CID 11920277.
Behre, Karl-Ernst (1999). "The history of beer additives in Europe - a review". Vegetation History and Archaeobotany. 8 (1–2): 35–48. Bibcode:1999VegHA...8...35B. doi:10.1007/BF02042841. S2CID 129429552.
Langezaal CR, Chandra A, Scheffer JJ (1992). "Antimicrobial screening of essential oils and extracts of some Humulus lupulus L. cultivars". Pharm Weekbl Sci. 14 (6): 353–356. doi:10.1007/bf01970171. PMID 1475174. S2CID 12561634.

theplantlist.org (Global: 139^th place; English: 1,390^th place)

"Humulus lupulus L.". World Checklist of Selected Plant Families. Royal Botanic Gardens, Kew. Retrieved 2 February 2016 – via The Plant List. Note that this website has been superseded by World Flora Online

tomsofmaine.com (Global: low place; English: low place)

"Hops [CO2] Extract". Toms of Maine. Archived from the original on December 24, 2013. Retrieved 2009-06-06.

umich.edu (Global: 429^th place; English: 307^th place)

climbers.lsa.umich.edu

Burnham, Robyn J. (2014). "Vitis riparia". CLIMBERS: Censusing Lianas in Mesic Biomes of Eastern Regions. University of Michigan College of Literature, Sciences and the Arts. Retrieved 2017-04-02.

usda.gov (Global: 512^th place; English: 322^nd place)

plants.sc.egov.usda.gov

NRCS. "Humulus lupulus var. neomexicanus". PLANTS Database. United States Department of Agriculture (USDA). Retrieved May 5, 2016.
NRCS. "Humulus lupulus var. pubescens". PLANTS Database. United States Department of Agriculture (USDA).

uwlax.edu (Global: low place; English: low place)

bioweb.uwlax.edu

Sewalish, Andrew. "Habitat & Adaptation". Humulus lupulus profile. Retrieved 2017-04-13.

web.archive.org (Global: 1^st place; English: 1^st place)

"Hops [CO2] Extract". Toms of Maine. Archived from the original on December 24, 2013. Retrieved 2009-06-06.
Plantlife website County Flowers page Archived 2015-04-30 at the Wayback Machine

wiley.com (Global: 150^th place; English: 174^th place)

onlinelibrary.wiley.com

Schönberger, C.; Kostelecky, T. (2011). "125th Anniversary Review: The Role of Hops in Brewing" (PDF). Journal of the Institute of Brewing. 117 (3): 259–267. doi:10.1002/j.2050-0416.2011.tb00471.x.

worldcat.org (Global: 4^th place; English: 4^th place)

search.worldcat.org

Eriksen, Renée L.; Padgitt-Cobb, Lillian K.; Townsend, M. Shaun; Henning, John A. (December 2021). "Gene expression for secondary metabolite biosynthesis in hop (Humulus lupulus L.) leaf lupulin glands exposed to heat and low-water stress". Scientific Reports. 11 (1): 5138. Bibcode:2021NatSR..11.5138E. doi:10.1038/s41598-021-84691-y. ISSN 2045-2322. PMC 7970847. PMID 33664420.
Goese, Markus; Kammhuber, Klaus; Bacher, Adelbert; Zenk, Meinhart H.; Eisenreich, Wolfgang (1999-07-15). "Biosynthesis of bitter acids in hops. A 13C-NMR and 2H-NMR study on the building blocks of humulone". European Journal of Biochemistry. 263 (2): 447–454. Bibcode:1999EJBio.263..447G. doi:10.1046/j.1432-1327.1999.00518.x. ISSN 0014-2956. PMID 10406953.
Natsume, S.; Takagi, H.; Shiraishi, A.; Murata, J.; Toyonaga, H.; Patzak, J.; Takagi, M.; Yaegashi, H.; Uemura, A.; Mitsuoka, C.; Yoshida, K. (2014-11-20). "The Draft Genome of Hop (Humulus lupulus), an Essence for Brewing". Plant and Cell Physiology. 56 (3): 428–441. doi:10.1093/pcp/pcu169. ISSN 0032-0781. PMID 25416290.
Padgitt-Cobb, Lillian K.; Kingan, Sarah B.; Wells, Jackson; Elser, Justin; Kronmiller, Brent; Moore, Daniel; Concepcion, Gregory; Peluso, Paul; Rank, David; Jaiswal, Pankaj; Henning, John; Hendrix, David A. (2021-02-18). "A draft phased assembly of the diploid Cascade hop ( Humulus lupulus ) genome". The Plant Genome. 14 (1) e20072. doi:10.1002/tpg2.20072. ISSN 1940-3372. PMID 33605092. S2CID 231962731.
Benkherouf, Ali Y.; Soini, Sanna L.; Stompor, Monika; Uusi-Oukari, Mikko (February 2019). "Positive allosteric modulation of native and recombinant GABAA receptors by hops prenylflavonoids". European Journal of Pharmacology. 852: 34–41. doi:10.1016/j.ejphar.2019.02.034. ISSN 0014-2999. PMID 30797788. S2CID 73456325.
Franco, L.; Sánchez, C.; Bravo, R.; Rodriguez, A.; Barriga, C.; Juánez, Javier Cubero (2012-06-01). "The sedative effects of hops (Humulus lupulus), a component of beer, on the activity/rest rhythm". Acta Physiologica Hungarica. 99 (2): 133–139. doi:10.1556/APhysiol.99.2012.2.6. ISSN 0231-424X. PMID 22849837.
Divashuk, Mikhail G.; Alexandrov, Oleg S.; Razumova, Olga V.; Kirov, Ilya V.; Karlov, Gennady I. (2014-01-21). "Molecular Cytogenetic Characterization of the Dioecious Cannabis sativa with an XY Chromosome Sex Determination System". PLOS ONE. 9 (1) e85118. Bibcode:2014PLoSO...985118D. doi:10.1371/journal.pone.0085118. ISSN 1932-6203. PMC 3897423. PMID 24465491.
Murakami, A; Darby, P; Javornik, B; Pais, M S S; Seigner, E; Lutz, A; Svoboda, P (2006-05-10). "Molecular phylogeny of wild Hops, Humulus lupulus L." Heredity. 97 (1): 66–74. Bibcode:2006Hered..97...66M. doi:10.1038/sj.hdy.6800839. ISSN 0018-067X. PMID 16685279. S2CID 11920277.
Pisupati, Rahul; Vergara, Daniela; Kane, Nolan C. (2018-02-21). "Diversity and evolution of the repetitive genomic content in Cannabis sativa". BMC Genomics. 19 (1): 156. doi:10.1186/s12864-018-4494-3. ISSN 1471-2164. PMC 5822635. PMID 29466945.
Laverty, Kaitlin U.; Stout, Jake M.; Sullivan, Mitchell J.; Shah, Hardik; Gill, Navdeep; Holbrook, Larry; Deikus, Gintaras; Sebra, Robert; Hughes, Timothy R.; Page, Jonathan E.; van Bakel, Harm (2018-11-08). "A physical and genetic map of Cannabis sativa identifies extensive rearrangements at the THC/CBD acid synthase loci". Genome Research. 29 (1): 146–156. doi:10.1101/gr.242594.118. ISSN 1088-9051. PMC 6314170. PMID 30409771.
Marks, M. David; Tian, Li; Wenger, Jonathan P.; Omburo, Stephanie N.; Soto-Fuentes, Wilfredo; He, Ji; Gang, David R.; Weiblen, George D.; Dixon, Richard A. (2009-07-06). "Identification of candidate genes affecting Δ9-tetrahydrocannabinol biosynthesis in Cannabis sativa". Journal of Experimental Botany. 60 (13): 3715–3726. doi:10.1093/jxb/erp210. ISSN 1460-2431. PMC 2736886. PMID 19581347.

Humulus lupulus (English Wikipedia)

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