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Koepfli, K.-P.; Jenks, S. M.; Eizirik, E.; Zahirpour, T.; Van Valkenburgh, B.; Wayne, R. K. (2006). "Molecular systematics of the Hyaenidae: Relationships of a relictual lineage resolved by a molecular supermatrix". Molecular Phylogenetics and Evolution. 38 (3): 603–620. Bibcode:2006MolPE..38..603K. doi:10.1016/j.ympev.2005.10.017. PMID16503281.
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