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Hochberg, M; Elmes, G. W.; Thomas, J. A.; Clarke, R. T (1996). "Mechanisms of local persistence in coupled host-parasitoid associations: the case model of Maculinea rebeli and Ichneumon eumerus". Philosophical Transactions of the Royal Society of London. Series B, Biological Sciences. 351 (1348): 1713–1724. Bibcode:1996RSPTB.351.1713H. doi:10.1098/rstb.1996.0153.