Manns U, Bremer B (2010). "Towards a better understanding of intertribal relationships and stable tribal delimitations within Cinchonoideae s.s. (Rubiaceae)". Molecular Phylogenetics and Evolution. 56 (1): 21–39. doi:10.1016/j.ympev.2010.04.002. PMID20382247.
Manns U, Bremer B (2010). "Towards a better understanding of intertribal relationships and stable tribal delimitations within Cinchonoideae s.s. (Rubiaceae)". Molecular Phylogenetics and Evolution. 56 (1): 21–39. doi:10.1016/j.ympev.2010.04.002. PMID20382247.