Matsuda, Keisuke; Adachi, Haruhiko; Gotoh, Hiroki; Inoue, Yasuhiro; Kondo, Shigeru (15 October 2024). "Adhesion and shrinkage transform the rounded pupal horn into an angular adult horn in Japanese rhinoceros beetle". Development. 151 (20). doi:10.1242/dev.202082. PMID38477641.
del Sol, Jillian F.; Hongo, Yoshihito; Boisseau, Romain P.; Berman, Gabriella H.; Allen, Cerisse E.; Emlen, Douglas. J. (February 2021). "Population differences in the strength of sexual selection match relative weapon size in the Japanese rhinoceros beetle, Trypoxylus dichotomus (Coleoptera: Scarabaeidae)†". Evolution. 75 (2): 394–413. doi:10.1111/evo.14101. PMID33009663.
Kojima, Wataru; Sugiura, Shinji; Makihara, Hiroshi; Ishikawa, Yukio; Takanashi, Takuma (1 March 2014). "Rhinoceros Beetles Suffer Male-Biased Predation by Mammalian and Avian Predators". Zoological Science. 31 (3): 109–115. doi:10.2108/zsj.31.109. PMID24601771.
Matsuda, Keisuke; Adachi, Haruhiko; Gotoh, Hiroki; Inoue, Yasuhiro; Kondo, Shigeru (15 October 2024). "Adhesion and shrinkage transform the rounded pupal horn into an angular adult horn in Japanese rhinoceros beetle". Development. 151 (20). doi:10.1242/dev.202082. PMID38477641.
del Sol, Jillian F.; Hongo, Yoshihito; Boisseau, Romain P.; Berman, Gabriella H.; Allen, Cerisse E.; Emlen, Douglas. J. (February 2021). "Population differences in the strength of sexual selection match relative weapon size in the Japanese rhinoceros beetle, Trypoxylus dichotomus (Coleoptera: Scarabaeidae)†". Evolution. 75 (2): 394–413. doi:10.1111/evo.14101. PMID33009663.
Kojima, Wataru; Sugiura, Shinji; Makihara, Hiroshi; Ishikawa, Yukio; Takanashi, Takuma (1 March 2014). "Rhinoceros Beetles Suffer Male-Biased Predation by Mammalian and Avian Predators". Zoological Science. 31 (3): 109–115. doi:10.2108/zsj.31.109. PMID24601771.