Leopard (English Wikipedia)

Allen, J. A. (1902). "Mammal names proposed by Oken in his 'Lehrbuch der Zoologie'". Bulletin of the American Museum of Natural History. 16 (27): 373−379. Archived from the original on 2023-12-09. Retrieved 2024-03-14.

archive.org (Global: 6^th place; English: 6^th place)

Lewis, C. T. & Short, C. (1879). "lěǒpardus". A Latin Dictionary. Oxford: Clarendon Press. p. 1069.
Liddell, H. G. & Scott, R. (1889). "λέο-πάρδος". A Greek–English Lexicon. Oxford: Clarendon Press. p. 884.
Partridge, E. (1983). Origins: A Short Etymological Dictionary of Modern English. New York: Greenwich House. p. 349. ISBN 978-0-517-41425-5.
Lewis, C. T. & Short, C. (1879). "panthera". A Latin Dictionary. Oxford: Clarendon Press. p. 1298.
Lewis, C. T. & Short, C. (1879). "pardus". A Latin Dictionary. Oxford: Clarendon Press. p. 1302.
Linnaeus, C. (1758). "Felis pardus". Caroli Linnæi Systema naturæ per regna tria naturæ, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Vol. Tomus I (decima, reformata ed.). Holmiae: Laurentius Salvius. pp. 41−42. (in Latin)
Ellerman, J. R.; Morrison-Scott, T. C. S. (1966). Checklist of Palaearctic and Indian mammals 1758 to 1946 (Second ed.). London: British Museum of Natural History. pp. 315–317.
Pocock, R. I. (1917). "The Classification of existing Felidae". The Annals and Magazine of Natural History. Series 8. XX: 329–350. doi:10.1080/00222931709487018.
Pocock, R. I. (1939). "Panthera pardus". The Fauna of British India, including Ceylon and Burma. Mammalia: Volume 1. London: Taylor and Francis. pp. 222–239.
Hemprich, W.; Ehrenberg, C. G. (1830). "Felis, pardus?, nimr". In Dr. C. G. Ehrenberg (ed.). Symbolae Physicae, seu Icones et Descriptiones Mammalium quae ex Itinere per Africam Borealem et Asiam Occidentalem Friderici Guilelmi Hemprich et Christiani Godofredi Ehrenberg. Decas Secunda. Zoologica I. Mammalia II. Berolini: Officina Academica. pp. Plate 17.
Gray, J. E. (1862). "Description of some new species of Mammalia". Proceedings of the Royal Zoological Society of London. 30: 261−263, plate XXXIII. doi:10.1111/j.1469-7998.1862.tb06524.x.
Schmid, E. (1940). "Variationstatistische Untersuchungen am Gebiss pleistozäner und rezenter Leoparden und anderer Feliden". Zeitschrift für Säugetierkunde. 15: 1–179.
Mivart, St. G. J. (1900). "Different kind of Cats". The Cat: An Introduction to the Study of Backboned Animals, Especially Mammals. London: John Murray. pp. 391–439.
Heptner, V. G. & Sludskii, A. A. (1992) [1972]. "Bars (leopard)". Mlekopitajuščie Sovetskogo Soiuza. Moskva: Vysšaia Škola [Mammals of the Soviet Union, Volume II, Part 2]. Washington DC: Smithsonian Institution and the National Science Foundation. pp. 203–273. ISBN 978-90-04-08876-4.
Divyabhanusinh (1993). "On mutant leopards Panthera pardus from India". Journal of the Bombay Natural History Society. 90 (1): 88−89.
Bertram, B. C. R. (1979). "Serengeti predators and their social systems". In Sinclair, A. R. E.; Norton-Griffiths, M. (eds.). Serengeti: dynamics of an ecosystem. University of Chicago Press. pp. 221–248.
Pedersen, C. F. (1971). The International Flag Book in Color. Morrow.
Anderson, K. (1954). "The Spotted Devil of Gummalapur". Nine Man-Eaters and one Rogue. London: George Allen & Unwin. pp. 36–51.

biodiversitylibrary.org (Global: 168^th place; English: 136^th place)

Prater, S. H. (1921). "Record panther skull (P. p. pardus)". The Journal of the Bombay Natural History Society. XXVII (Part IV): 933–935. Archived from the original on 2019-05-02. Retrieved 2019-08-24.

biosoil.ru (Global: low place; English: low place)

Uphyrkina, O.; Johnson, E. W.; Quigley, H.; Miquelle, D.; Marker, L.; Bush, M. & O'Brien, S. J. (2001). "Phylogenetics, genome diversity and origin of modern leopard, Panthera pardus" (PDF). Molecular Ecology. 10 (11): 2617–2633. Bibcode:2001MolEc..10.2617U. doi:10.1046/j.0962-1083.2001.01350.x. PMID 11883877. S2CID 304770. Archived (PDF) from the original on 2011-09-10.

books.google.com (Global: 3^rd place; English: 3^rd place)

Oken, L. (1816). "1. Art, Panthera". Lehrbuch der Zoologie. 2. Abtheilung. Jena: August Schmid & Comp. p. 1052. Archived from the original on 2024-05-05. Retrieved 2019-04-06.
Meyer, F. A. A. (1794). "Über de la Metheries schwarzen Panther". Zoologische Annalen. Erster Band. Weimar: Im Verlage des Industrie-Comptoirs. pp. 394–396. Archived from the original on 2024-05-05. Retrieved 2019-04-05.
Kawata, K. (2001). "Zoological gardens of Japan". In Kisling, V.N. (ed.). Zoo and Aquarium History: Ancient Animal Collections to Zoological Gardens. Boca Raton, Florida: CRC Press. pp. 295–329. ISBN 978-0-8493-2100-9. Archived from the original on 2024-05-05. Retrieved 2021-09-07.
Mills, M. G. L. (2005). "Subfamily Pantherinae". In Skinner, J. D.; Chimimba, C. T. (eds.). The mammals of the southern African sub region (Third ed.). Cambridge: Cambridge University Press. pp. 385–396. ISBN 978-0-521-84418-5. Archived from the original on 2024-05-05. Retrieved 2021-05-19.
Hoath, R. (2009). "Leopard Panthera pardus (Linnaeus, 1758)". Field Guide to the Mammals of Egypt. Cairo, Egypt: American University in Cairo Press. pp. 106–107. ISBN 978-977-416-254-1. Archived from the original on 2024-05-05. Retrieved 2021-05-19.
Estes, R. (1991). "Leopard Panthera pardus". The Behavior Guide to African Mammals, Including Hoofed Mammals, Carnivores, Primates. Los Angeles: The University of California Press. pp. 366–369. ISBN 978-0-520-08085-0. Archived from the original on 2024-05-05. Retrieved 2021-05-19.
Hunter, L.; Henschel, P. & Ray, J. C. (2013). "Panthera pardus Leopard". In Kingdon, J.; Happold, D.; Butynski, T.; Hoffmann, M.; Happold, M. & Kalina, J. (eds.). Mammals of Africa. London: Bloomsbury Publishing. pp. 159–168. ISBN 978-1-4081-8996-2. Archived from the original on 2024-05-05. Retrieved 2021-05-19.
Nowak, R. M. (1999). "Panthera pardus (Leopard)". Walker's Mammals of the World (Sixth ed.). Baltimore, US: Johns Hopkins University Press. pp. 828–831. ISBN 978-0-8018-5789-8. Archived from the original on 2024-05-05. Retrieved 2021-05-19.
Sunquist, M. E. & Sunquist, F. (2002). "Leopard Panthera pardus". Wild Cats of the World. Chicago: University of Chicago Press. pp. 318–342. ISBN 978-0-226-77999-7. Archived from the original on 2024-04-30. Retrieved 2021-05-19.
Bothma, Jacobus du P.; Walker, Clive (2013-11-11). Larger Carnivores of the African Savannas. Springer Science & Business Media. ISBN 978-3-662-03766-9. Archived from the original on 2024-01-06. Retrieved 2024-01-06.
Schaller, G. (1972). Serengeti: a kingdom of predators. New York: Knopf. ISBN 978-0-394-47242-3. Archived from the original on 2024-05-05. Retrieved 2020-09-18.
Haist, M. (1999). "The Lion, bloodline, and kingship". In Hassig, D. (ed.). The Mark of the Beast: The Medieval Bestiary in Art, Life, and Literature. London: Taylor & Francis. pp. 3–16. ISBN 978-0-8153-2952-7. Archived from the original on 2024-05-05. Retrieved 2021-09-07.

brainmuseum.org (Global: low place; English: low place)

"Leopard (Panthera pardus); Physical characteristics and distribution". Comparative Mammalian Brain Collections. Archived from the original on 2011-01-03. Retrieved 2008-06-07.

britishmuseum.org (Global: 1,421^st place; English: 1,087^th place)

"Benin: an African kingdom" (PDF). London: British Museum. Archived (PDF) from the original on 2008-08-05. Retrieved 2016-03-29.

bucknell.edu (Global: 671^st place; English: 1,144^th place)

departments.bucknell.edu

Wozencraft, W. C. (2005). "Species Panthera pardus". In Wilson, D. E.; Reeder, D. M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Johns Hopkins University Press. p. 547. ISBN 978-0-8018-8221-0. OCLC 62265494.

cambridge.org (Global: 209^th place; English: 151^st place)

Marciszak, A.; Lipecki, G.; Gornig, W.; Matyaszczyk, L.; Oszczepalińska, O.; Nowakowski, D.; Talamo, S. (2022). "The first radiocarbon-dated remains of the Leopard Panthera pardus (Linnaeus, 1758) from the Pleistocene of Poland". Radiocarbon. 64 (6): 1359–1372. Bibcode:2022Radcb..64.1359M. doi:10.1017/RDC.2022.33. hdl:11585/887180. S2CID 255850069. Archived from the original on 2024-05-05. Retrieved 2023-07-18.

catsg.org (Global: low place; English: low place)

Spalton, J. A. & Al Hikmani, H. M. (2006). "The Leopard in the Arabian Peninsula – Distribution and subspecies status" (PDF). Cat News (Special Issue 1): 4–8. Archived (PDF) from the original on 2015-06-19.
Spalton, J. A. & Al Hikmani, H. M. (2006). "The Leopard in the Arabian Peninsula – Distribution and subspecies status" (PDF). Cat News (Special Issue 1): 4–8.
Judas, J.; Paillat, P.; Khoja, A. & Boug, A. (2006). "Status of the Arabian leopard in Saudi Arabia" (PDF). Cat News (Special Issue 1): 11–19. Archived from the original (PDF) on 2015-09-19. Retrieved 2019-03-17.
Al Jumaily, M.; Mallon, D. P.; Nasher, A. K. & Thowabeh, N. (2006). "Status Report on Arabian Leopard in Yemen" (PDF). Cat News (Special Issue 1): 20–25.
Rozhnov, V.V.; Yachmennikova, A.A.; Dronova, N.A.; Naidenko, S.V.; Hernandez-Blanco, J.A.; Chistopolova, M.D.; Pkhitikov, A.B.; Tembotova, F.A.; Trepet, S.A. & Chestin, I.E. (2022). "Experience of the leopard re-covering through reintroduction in the Russian Caucasus" (PDF). Cat News (Special issue): 67–71. Archived (PDF) from the original on 2023-11-11. Retrieved 2023-11-11.

conservationindia.org (Global: low place; English: low place)

Arthreya, V. (2012). "Living with Leopards Outside Protected Areas in India". Conservation India. Archived from the original on 2012-12-21. Retrieved 2012-12-10.

csulb.edu (Global: low place; English: low place)

web.csulb.edu

Davis, B. W.; Li, G. & Murphy, W. J. (2010). "Supermatrix and species tree methods resolve phylogenetic relationships within the big cats, Panthera (Carnivora: Felidae)" (PDF). Molecular Phylogenetics and Evolution. 56 (1): 64–76. Bibcode:2010MolPE..56...64D. doi:10.1016/j.ympev.2010.01.036. PMID 20138224. Archived from the original (PDF) on 2016-03-05.

cuny.edu (Global: 2,325^th place; English: 1,345^th place)

geo.hunter.cuny.edu

Murphey, R. (1951). "The Decline of North Africa Since the Roman Occupation: Climatic or Human?" (PDF). Annals of the Association of American Geographers. XLI (2): 116–132. doi:10.1080/00045605109352048. Archived (PDF) from the original on 2006-09-14.

doi.org (Global: 2^nd place; English: 2^nd place)

Stein, A.B.; Gerngross, P.; Al Hikmani, H.; Balme, G.; Bertola, L.; Drouilly, M.; Farhadinia, M.S.; Feng, L.; Ghoddousi, A.; Henschel, P.; Jhala, Y.; Khorozyan, I.; Kittle, A.; Laguardia, A.; Luo, S.-J.; Mann, G.; Miquelle, D.; Moheb, Z.; Raza, H.; Rostro-García, S.; Shivakumar, S.; Song, D. & Wibisono, H. (2025). "Panthera pardus". IUCN Red List of Threatened Species. 2025 e.T15954A286153337. doi:10.2305/IUCN.UK.2025-2.RLTS.T15954A286153337.en.
Pocock, R. I. (1917). "The Classification of existing Felidae". The Annals and Magazine of Natural History. Series 8. XX: 329–350. doi:10.1080/00222931709487018.
Miththapala, S.; Seidensticker, J. & O'Brien, S. J. (1996). "Phylogeographic subspecies recognition in leopards (Panthera pardus): molecular genetic variation" (PDF). Conservation Biology. 10 (4): 1115–1132. Bibcode:1996ConBi..10.1115M. doi:10.1046/j.1523-1739.1996.10041115.x. Archived (PDF) from the original on 2016-02-17. Retrieved 2017-03-09.
Uphyrkina, O.; Johnson, E. W.; Quigley, H.; Miquelle, D.; Marker, L.; Bush, M. & O'Brien, S. J. (2001). "Phylogenetics, genome diversity and origin of modern leopard, Panthera pardus" (PDF). Molecular Ecology. 10 (11): 2617–2633. Bibcode:2001MolEc..10.2617U. doi:10.1046/j.0962-1083.2001.01350.x. PMID 11883877. S2CID 304770. Archived (PDF) from the original on 2011-09-10.
Laguardia, A.; Kamler, J. F.; Li, S.; Zhang, C.; Zhou, Z. & Shi, K. (2017). "The current distribution and status of leopards Panthera pardus in China". Oryx. 51 (1): 153−159. doi:10.1017/S0030605315000988.
Wibisono, H.; Wilianto, E.; Pinondang, I.; Rahman, D.A. & Chandradewi, D. (2021). "Panthera pardus ssp. melas". IUCN Red List of Threatened Species. 2021 e.T15962A50660931. doi:10.2305/IUCN.UK.2021-2.RLTS.T15962A50660931.en. Retrieved 11 November 2023.
Ghoddousi, A. & Khorozyan, I. (2023). "Panthera pardus ssp. tulliana". IUCN Red List of Threatened Species. 2023 e.T15961A50660903. doi:10.2305/IUCN.UK.2023-1.RLTS.T15961A50660903.en. Retrieved 17 March 2024.
Khorozyan, I. G.; Gennady, F.; Baryshnikov, G. F. & Abramov, A. V. (2006). "Taxonomic status of the leopard, Panthera pardus (Carnivora, Felidae) in the Caucasus and adjacent areas". Russian Journal of Theriology. 5 (1): 41–52. doi:10.15298/rusjtheriol.05.1.06.
Gray, J. E. (1862). "Description of some new species of Mammalia". Proceedings of the Royal Zoological Society of London. 30: 261−263, plate XXXIII. doi:10.1111/j.1469-7998.1862.tb06524.x.
Anco, C.; Kolokotronis, S. O.; Henschel, P.; Cunningham, S. W.; Amato, G. & Hekkala, E. (2017). "Historical mitochondrial diversity in African leopards (Panthera pardus) revealed by archival museum specimens". Mitochondrial DNA Part A. 29 (3): 455–473. doi:10.1080/24701394.2017.1307973. PMID 28423965. S2CID 4348541.
Johnson, W. E.; Eizirik, E.; Pecon-Slattery, J.; Murphy, W. J.; Antunes, A.; Teeling, E. & O'Brien, S. J. (2006). "The late Miocene radiation of modern Felidae: a genetic assessment". Science. 311 (5757): 73–77. Bibcode:2006Sci...311...73J. doi:10.1126/science.1122277. PMID 16400146. S2CID 41672825. Archived from the original on 2020-10-04. Retrieved 2019-07-05.
Davis, B. W.; Li, G. & Murphy, W. J. (2010). "Supermatrix and species tree methods resolve phylogenetic relationships within the big cats, Panthera (Carnivora: Felidae)" (PDF). Molecular Phylogenetics and Evolution. 56 (1): 64–76. Bibcode:2010MolPE..56...64D. doi:10.1016/j.ympev.2010.01.036. PMID 20138224. Archived from the original (PDF) on 2016-03-05.
Mazák, J. H.; Christiansen, P.; Kitchener, A. C. & Goswami, A. (2011). "Oldest known pantherine skull and evolution of the tiger". PLOS ONE. 6 (10) e25483. Bibcode:2011PLoSO...625483M. doi:10.1371/journal.pone.0025483. PMC 3189913. PMID 22016768.
Bininda-Emonds, O. R. P.; Decker-Flum, D. M. & Gittleman, J. L. (2001). "The utility of chemical signals as phylogenetic characters: an example from the Felidae". Biological Journal of the Linnean Society. 72 (1): 1–15. doi:10.1111/j.1095-8312.2001.tb01297.x.
Tseng, Z. J.; Wang, X.; Slater, G. J.; Takeuchi, G. T.; Li, Q.; Liu, J. & Xie, G. (2014). "Himalayan fossils of the oldest known pantherine establish ancient origin of big cats". Proceedings of the Royal Society B: Biological Sciences. 281 (1774) 20132686. Bibcode:2014PBioS.28132686T. doi:10.1098/rspb.2013.2686. PMC 3843846. PMID 24225466.
Li, G.; Davis, B. W.; Eizirik, E. & Murphy, W. J. (2016). "Phylogenomic evidence for ancient hybridization in the genomes of living cats (Felidae)". Genome Research. 26 (1): 1–11. doi:10.1101/gr.186668.114. PMC 4691742. PMID 26518481.
Paijmans, J. L. A.; Barlow, A.; Förster, D. W.; Henneberger, K.; Meyer, M.; Nickel, B.; Nagel, D.; Havmøller, R. W.; Baryshnikov, G. F.; Joger, U.; Rosendahl, W. & Hofreiter, M. (2018). "Historical biogeography of the leopard (Panthera pardus) and its extinct Eurasian populations". BMC Evolutionary Biology. 18 (1): 156. Bibcode:2018BMCEE..18..156P. doi:10.1186/s12862-018-1268-0. PMC 6198532. PMID 30348080.
Marciszak, A.; Lipecki, G.; Gornig, W.; Matyaszczyk, L.; Oszczepalińska, O.; Nowakowski, D.; Talamo, S. (2022). "The first radiocarbon-dated remains of the Leopard Panthera pardus (Linnaeus, 1758) from the Pleistocene of Poland". Radiocarbon. 64 (6): 1359–1372. Bibcode:2022Radcb..64.1359M. doi:10.1017/RDC.2022.33. hdl:11585/887180. S2CID 255850069. Archived from the original on 2024-05-05. Retrieved 2023-07-18.
Diedrich, C. G. (2013). "Late Pleistocene leopards across Europe – northernmost European German population, highest elevated records in the Swiss Alps, complete skeletons in the Bosnia Herzegowina Dinarids and comparison to the Ice Age cave art". Quaternary Science Reviews. 76: 167–193. Bibcode:2013QSRv...76..167D. doi:10.1016/j.quascirev.2013.05.009.
Sauqué, V. & Cuenca-Bescós, G. (2013). "The Iberian Peninsula, the last European refugium of Panthera pardus Linnaeus 1758 during the Upper Pleistocene". Quaternaire. 24 (1): 13–24. doi:10.4000/quaternaire.6468. Archived from the original on 2023-07-18. Retrieved 2023-07-18.
Marciszak, A. & Stefaniak, K. (2010). "Two forms of cave lion: Middle Pleistocene Panthera spelaea fossilis Reichenau, 1906 and Upper Pleistocene Panthera spelaea spelaea Goldfuss, 1810 from the Bísnik Cave, Poland". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 258 (3): 339–351. Bibcode:2010NJGPA.258..339M. doi:10.1127/0077-7749/2010/0117. Archived from the original on 2018-09-25. Retrieved 2017-08-24.
Sauqué, V.; Rabal-Garcés, R. & Cuenca-Bescós, G. (2016). "Carnivores from Los Rincones, a leopard den in the highest mountain of the Iberian range (Moncayo, Zaragoza, Spain)". Historical Biology. 28 (4): 479–506. Bibcode:2016HBio...28..479S. doi:10.1080/08912963.2014.977882.
Sanchis, A.; Real, C.; Sauqué, V.; Núñez-Lahuerta, C.; Égüez, N.; Tormo, C.; Ripoll, M. P.; Carrión, Y. M.; Duarte, E. & de la Rasilla, M. (2019). "Neanderthal and carnivore activities at Llonin Cave, Asturias, northern Iberian Peninsula: Faunal study of Mousterian levels (MIS 3)". Comptes Rendus Palevol. 18 (1): 113–141. Bibcode:2019CRPal..18..113S. doi:10.1016/j.crpv.2018.06.001. hdl:10651/49880. S2CID 134157124.
Ghezzo, E. & Rook, L. (2015). "The remarkable Panthera pardus (Felidae, Mammalia) record from Equi (Massa, Italy): taphonomy, morphology, and paleoecology". Quaternary Science Reviews. 110: 131–151. doi:10.1016/j.quascirev.2014.12.020.
Mecozzi, Beniamino; Sardella, Raffaele; Boscaini, Alberto; Cherin, Marco; Costeur, Loïc; Madurell-Malapeira, Joan; Pavia, Marco; Profico, Antonio; Iurino, Dawid A. (15 June 2021). "The tale of a short-tailed cat: New outstanding Late Pleistocene fossils of Lynx pardinus from southern Italy". Quaternary Science Reviews. 262 106840. Bibcode:2021QSRv..26206840M. doi:10.1016/j.quascirev.2021.106840. hdl:2434/959858.
Sommer, R. S. & Benecke, N. (2006). "Late Pleistocene and Holocene development of the felid fauna (Felidae) of Europe: a review". Journal of Zoology. 269 (1): 7–19. Bibcode:2006JZoo..269....7S. doi:10.1111/j.1469-7998.2005.00040.x.
Miracle, P. T.; Lenardić, J. M.; Brajković, D. (2010). "Last glacial climates, "Refugia", and faunal change in Southeastern Europe: Mammalian assemblages from Veternica, Velika pećina, and Vindija caves (Croatia)". Quaternary International. 212 (2): 137–148. Bibcode:2010QuInt.212..137M. doi:10.1016/j.quaint.2009.06.003.
Chi T.-C.; Gan Y.; Yang T.-R. & Chang, C.-H. (2021). "First report of leopard fossils from a limestone cave in Kenting area, southern Taiwan". PeerJ. 9 e12020. doi:10.7717/peerj.12020. PMC 8388558. PMID 34513335.
Sun, X.; Peng, L.; Tsutaya, T.; Jiangzuo, Q.; Hasegawa, Y.; Hou, Y.; Han, Y.; Zhuang, Y.; Ramos Madrigal, J.; Taurozzi, A. J.; Mackie, M.; Trochė, G.; Olsen, J. V.; Cappellini, E.; O'Brien, S. J.; Gilbert, M. T. P.; Yamaguchi, N.; Luo, S.-J. (2026). "The Japanese Archipelago sheltered cave lions, not tigers, during the Late Pleistocene". Proceedings of the National Academy of Sciences of the United States of America. 123 (6) e2523901123. Bibcode:2026PNAS..12323901S. doi:10.1073/pnas.2523901123. PMC 12890994. PMID 41587328.
Pocook, R. I. (1932). "The Leopards of Africa". Proceedings of the Zoological Society of London. 102 (2): 543–591. doi:10.1111/j.1096-3642.1932.tb01085.x.
Allen, W. L.; Cuthill, I. C.; Scott-Samuel, N. E. & Baddeley, R. (2010). "Why the leopard got its spots: relating pattern development to ecology in felids". Proceedings of the Royal Society B. 278 (1710): 1373–1380. doi:10.1098/rspb.2010.1734. PMC 3061134. PMID 20961899.
Stein, A. B. & Hayssen, V. (2010). "Panthera pardus (Carnivora: Felidae)". Mammalian Species. 45 (900): 30–48. doi:10.1644/900.1. PMC 7149468. S2CID 44839740.
Robinson, R. (1970). "Inheritance of the black form of the leopard Panthera pardus". Genetica. 41 (1): 190–197. doi:10.1007/BF00958904. PMID 5480762. S2CID 5446868.
Eizirik, E.; Yuhki, N.; Johnson, W. E.; Menotti-Raymond, M.; Hannah, S. S.; O'Brien, S. J. (2003). "Molecular genetics and evolution of melanism in the cat family". Current Biology. 13 (5): 448–453. Bibcode:2003CBio...13..448E. doi:10.1016/S0960-9822(03)00128-3. PMID 12620197. S2CID 19021807.
Kawanishi, K.; Sunquist, M. E.; Eizirik, E.; Lynam, A. J.; Ngoprasert, D.; Wan Shahruddin, W. N.; Rayan, D. M.; Sharma, D. S. K. & Steinmetz, R. (2010). "Near fixation of melanism in leopards of the Malay Peninsula". Journal of Zoology. 282 (3): 201–206. doi:10.1111/j.1469-7998.2010.00731.x.
Da Silva L. G., K.; Kawanishi, K.; Henschel P.; Kittle, A.; Sanei, A.; Reebin, A.; Miquelle, D.; Stein, A. B.; Watson, A.; Kekule, L. B.; Machado, R. B. & Eizirik, E. (2017). "Mapping black panthers: Macroecological modeling of melanism in leopards (Panthera pardus)". PLOS ONE. 12 (4) e0170378. Bibcode:2017PLoSO..1270378D. doi:10.1371/journal.pone.0170378. PMC 5381760. PMID 28379961.
Pirie, T. J.; Thomas, R. L. & Fellowes, M. D. E. (2016). "Erythristic leopards Panthera pardus in South Africa". Bothalia. 46 (1): 1–5. doi:10.4102/abc.v46i1.2034.
Powell, J.; Axmacher, J. C.; Linnell, J. D.C. & Durant, S. M. (2021). "Diverse locations and a long history: Historical context for urban Leopards (Panthera pardus) in the early anthropocene From Seoul, Korea". Frontiers in Conservation Science. 2 765911. Bibcode:2021FrCS....2.5911P. doi:10.3389/fcosc.2021.765911. hdl:11250/2999764.
Jacobson, A. P.; Gerngross, P.; Lemeris, J. R. Jr.; Schoonover, R. F.; Anco, C.; Breitenmoser-Würsten, C.; Durant, S. M.; Farhadinia, M. S.; Henschel, P.; Kamler, J. F.; Laguardia, A.; Rostro-García, S.; Stein, A. B. & Dollar, L. (2016). "Leopard (Panthera pardus) status, distribution, and the research efforts across its range". PeerJ. 4 e1974. doi:10.7717/peerj.1974. PMC 4861552. PMID 27168983.
Pirie, T. J.; Thomas, R. L. & Fellowes, M. D. E. (2017). "Increasing game prices may alter farmers' behaviours towards leopards (Panthera pardus) and other carnivores in South Africa". PeerJ. 5 e3369. doi:10.7717/peerj.3369. PMC 5452990. PMID 28584709.
Soultan, A.; Attum, O.; Hamada, A.; Hatab, E. B.; Ahmed, S. E.; Eisa, A.; Al Sharif, I.; Nagy, A. & Shohdi, W. (2017). "Recent observation for leopard Panthera pardus in Egypt". Mammalia. 81 (1): 115–117. doi:10.1515/mammalia-2015-0089. S2CID 90676105.
Karataş, A.; Bulut, Ş. & Akbaba, B. (2021). "Camera trap records confirm the survival of the Leopard (Panthera pardus L., 1758) in eastern Turkey (Mammalia: Felidae)". Zoology in the Middle East. 67 (3): 198–205. doi:10.1080/09397140.2021.1924419. hdl:11491/7382. S2CID 235564429.
Ünal, Y.; Uysal, H.; Koca, A. & Zenbilci, M. (2023). "New records of the living Anatolian Leopard (Panthera pardus tulliana L., 1758) in the Mediterranean region of Turkey". Applied Ecology and Environmental Research. 21 (2): 1043–1059. Bibcode:2023ApEER..21.1043U. doi:10.15666/aeer/2102_10431059.
Asad, M.; Waseem, M.; Ross, J. G. & Paterson, A. M. (2019). "The Un-Common Leopard: presence, distribution and abundance in Gallies and Murree Forest Division, Northern Pakistan". Nature Conservation. 37: 53–80. Bibcode:2019NaCon..37...53A. doi:10.3897/natureconservation.37.32748. hdl:10182/11138.
Akrim, F.; Kellner, K. F.; Mahmood, T.; Waseem, M.; Zangi, I.-U.-D.; Qasim, S.; Aslam, A.; Hashmi, S. A.; Shoukat, H. B.; Raqeeb, M. & Belant, J. L. (2025). "Habitat suitability of common leopard in northern Pakistan". Ecosphere. 16 (3) e70218. Bibcode:2025Ecosp..16E0218A. doi:10.1002/ecs2.70218.
Kittle, A. M.; Watson, A. C.; Kumara, P. H. S. C.; Sandanayake, S. D. K. C.; Sanjeewani, H. K. N. & Fernando, T. S. P. (2014). "Notes on the diet and habitat selection of the Sri Lankan Leopard Panthera pardus kotiya (Mammalia: Felidae) in the central highlands of Sri Lanka". Journal of Threatened Taxa. 6 (9): 6214–6221. doi:10.11609/JoTT.o3731.6214-21.
Saw Sha Bwe Moo; Froese, G.Z.L. & Gray, T.N.E. (2017). "First structured camera-trap surveys in Karen State, Myanmar, reveal high diversity of globally threatened mammals". Oryx. 52 (3): 537−543. doi:10.1017/S0030605316001113.
Rostro-García, S.; Kamler, J. F.; Ash, E.; Clements, G. R.; Gibson, L.; Lynam, A. J.; McEwin, R.; Naing, H. & Paglia, S. (2016). "Endangered leopards: Range collapse of the Indochinese leopard (Panthera pardus delacouri) in Southeast Asia". Biological Conservation. 201: 293–300. Bibcode:2016BCons.201..293R. doi:10.1016/j.biocon.2016.07.001. hdl:10722/232870.
Johnson, A.; Vongkhamheng, C.; Hedemark, M. & Saithongdam, T. (2006). "Effects of human–carnivore conflict on tiger (Panthera tigris) and prey populations in Lao PDR" (PDF). Animal Conservation. 9 (4): 421–430. Bibcode:2006AnCon...9..421J. doi:10.1111/j.1469-1795.2006.00049.x. S2CID 73637721. Archived (PDF) from the original on 2017-08-10.
Wibisono, H. T.; Wahyudi, H. A.; Wilianto, E.; Pinondang, I. M. R.; Primajati, M.; Liswanto, D. & Linkie, M. (2018). "Identifying priority conservation landscapes and actions for the Critically Endangered Javan leopard in Indonesia: Conserving the last large carnivore in Java Island". PLOS ONE. 13 (6) e0198369. Bibcode:2018PLoSO..1398369W. doi:10.1371/journal.pone.0198369. PMC 6021038. PMID 29949588.
Growcott, J; Lobora, A; Markham, A; Searle, C. E.; Wahlström, J; Wijers, M; Simmons, B. I. (2024). "The secret acoustic world of leopards: A paired camera trap and bioacoustics survey facilitates the individual identification of leopards via their roars". Remote Sensing in Ecology and Conservation. 11 (4): 361–374. doi:10.1002/rse2.429.
Ortolani, A. (1999). "Spots, stripes, tail tips and dark eyes: predicting the function of carnivore colour patterns using the comparative method". Biological Journal of the Linnean Society. 67 (4): 433–476. doi:10.1111/j.1095-8312.1999.tb01942.x.
Caro, T. (2005). "The adaptive significance of coloration in mammals". BioScience. 55 (2): 125–136. doi:10.1641/0006-3568(2005)055[0125:TASOCI]2.0.CO;2.
Spalton, J.A.; Al Hikmani, H. M.; Willis, D. & Said, A. S. B. (2006). "Critically endangered Arabian leopards Panthera pardus nimr persist in the Jabal Samhan Nature Reserve, Oman". Oryx. 40 (3): 287–294. Bibcode:2006Oryx...40..287S. doi:10.1017/S0030605306000743.
Jenny, D. & Zuberbuhler, K. (2005). "Hunting behaviour in west African forest leopards". African Journal of Ecology. 43 (3): 197–200. Bibcode:2005AfJEc..43..197J. doi:10.1111/j.1365-2028.2005.00565.x. Archived from the original on 2018-09-25. Retrieved 2018-09-25.
Pirie, T. J.; Thomas, R. L.; Reilly, B. K. & Fellowes, M. D. E. (2014). "Social interactions between a male leopard (Panthera pardus) and two generations of his offspring". African Journal of Ecology. 52 (4): 574–576. Bibcode:2014AfJEc..52..574P. doi:10.1111/aje.12154.
Jenny, D. (1996). "Spatial organization of leopards Panthera pardus in Tai National Park, Ivory Coast: Is rainforest habitat a "tropical haven"?". Journal of Zoology. 240 (3): 427–440. Bibcode:1996JZoo..240..427J. doi:10.1111/j.1469-7998.1996.tb05296.x. Archived from the original on 2024-05-05. Retrieved 2021-09-06.
Mizutani, F. & Jewell, P. A. (1998). "Home-range and movements of leopards (Panthera pardus) on a livestock ranch in Kenya". Journal of Zoology. 244 (2): 269–286. doi:10.1017/S0952836998002118.
Stander, P. E.; Haden, P. J.; Kaqece, II. & Ghau, II. (1997). "The ecology of asociality in Namibian leopards". Journal of Zoology. 242 (2): 343–364. doi:10.1111/j.1469-7998.1997.tb05806.x.
Odden, M. & Wegge, P. (2005). "Spacing and activity patterns of leopards Panthera pardus in the Royal Bardia National Park, Nepal". Wildlife Biology. 11 (2): 145–152. doi:10.2981/0909-6396(2005)11[145:SAAPOL]2.0.CO;2. S2CID 86140708.
Hayward, M.W.; Henschel, P.; O'Brien, J.; Hofmeyr, M.; Balme, G. & Kerley, G. I. H. (2006). "Prey preferences of the leopard (Panthera pardus)" (PDF). Journal of Zoology. 270 (4): 298–313. doi:10.1111/j.1469-7998.2006.00139.x. Archived (PDF) from the original on 2012-11-05.
Johnson, K. G.; Wei, W.; Reid, D. G.; Jinchu, H. (1993). "Food habits of Asiatic leopards (Panthera pardus fusca) in Wolong Reserve, Sichuan, China". Journal of Mammalogy. 74 (3): 646–650. Bibcode:1993JMamm..74..646J. doi:10.2307/1382285. JSTOR 1382285.
Kruuk, H. & Turner, M. (1967). "Comparative notes on predation by lion, leopard, cheetah and wild dog in the Serengeti area, East Africa". Mammalia. 31 (1): 1–27. doi:10.1515/mamm.1967.31.1.1. S2CID 84619500.
Sauqué, Víctor; Rabal-Garcés, Raquel; Sola-Almagro, Cristina; Cuenca-Bescós, Gloria (18 March 2014). Rook, Lorenzo (ed.). "Bone Accumulation by Leopards in the Late Pleistocene in the Moncayo Massif (Zaragoza, NE Spain)". PLoS ONE. 9 (3) e92144. Bibcode:2014PLoSO...992144S. doi:10.1371/journal.pone.0092144. ISSN 1932-6203. PMC 3958443. PMID 24642667.
Blanco, R. E.; Luzardo-Vera, D. E. (2021). "A biomechanical model to assess the injury risk of leopards (Panthera pardus) hunting by free falling from trees". Biological Communications. 66 (3). doi:10.21638/spbu03.2021.305. hdl:20.500.12008/38512. ISSN 2587-5779.
Rafiq, K.; Wayward, M. W.; Wilson, A. M.; Meloro, C.; Jordan, N. R.; Wich, S. A.; McNutt, J. W.; Golabek, K. A. (2020). "Spatial and temporal overlaps between leopards (Panthera pardus) and their competitors in the African large predator guild". Journal of Zoology. 311 (4): 246–259. doi:10.1111/jzo.12781. hdl:1959.4/unsworks_67550.
Balme, G. A.; Pitman, R. T.; Robinson, H. S.; Miller, J. R. B.; Funston, P. J.; Hunter, L. T. B. (2017). "Leopard distribution and abundance is unaffected by interference competition with lions". Behavioral Ecology. 28 (5): 1348–1358. doi:10.1093/beheco/arx098.
Balme, G. A.; Miller, J. R. B.; Pitman, R. T.; Hunter, L. T. B. (2017). "Caching reduces kleptoparasitism in a solitary, large felid". Journal of Animal Ecology. 86 (3): 634–644. Bibcode:2017JAnEc..86..634B. doi:10.1111/1365-2656.12654. PMID 28217865.
Vissia, S.; Fattebert, J.; van Langevelde, F. (2022). "Leopard density and interspecific spatiotemporal interactions in a hyena-dominated landscape". Ecology and Evolution. 12 (10) e9365. Bibcode:2022EcoEv..12E9365V. doi:10.1002/ece3.9365. PMC 9534747. PMID 36225822.
Miller, J. R. B.; Pitman, R. T.; Mann, G. K. H.; Fuller, A. K.; Balme, G. A. (2018). "Lions and leopards coexist without spatial, temporal or demographic effects of interspecific competition". Journal of Animal Ecology. 87 (6): 1709–1726. Bibcode:2018JAnEc..87.1709M. doi:10.1111/1365-2656.12883. PMID 30010193.
Srivathsa, A.; Ramachandran, V.; Saravanan, P.; Sureshbabu, A.; Ganguly, D. & Ramakrishnan, U. (2023). "Topcats and underdogs: intraguild interactions among three apex carnivores across Asia's forestscapes". Biological Reviews. 98 (6): 2114–2135. doi:10.1111/brv.12998. PMID 37449566. S2CID 259903849.
Thinley, P.; Rajaratnam, R.; Lassoie, J. P.; Morreale, S. J.; Curtis, P. D.; Vernes, K.; Leki Leki; Phuntsho, S.; Dorji, T. & Dorji, P. (2018). "The ecological benefit of tigers (Panthera tigris) to farmers in reducing crop and livestock losses in the eastern Himalayas: Implications for conservation of large apex predators". Biological Conservation. 219: 119–125. doi:10.1016/j.biocon.2018.08.007.
Karanth, K. U. & Sunquist, M. E. (2000). "Behavioural correlates of predation by tiger (Panthera tigris), leopard (Panthera pardus) and dhole (Cuon alpinus) in Nagarahole, India". Journal of Zoology. 250 (2): 255–265. doi:10.1111/j.1469-7998.2000.tb01076.x.
Karanth, K. U. & Sunquist, M. E. (1995). "Prey selection by tiger, leopard and dhole in tropical forests". Journal of Animal Ecology. 64 (4): 439–450. Bibcode:1995JAnEc..64..439K. doi:10.2307/5647. JSTOR 5647.
Steinmetz, R.; Seuaturien, N.; Intanajitjuy, P.; Inrueang, P. & Prempree, K. (2021). "The effects of prey depletion on dietary niches of sympatric apex predators in Southeast Asia". Integrative Zoology. 16 (1): 19–32. doi:10.1111/1749-4877.12461. PMID 32627329.
Odden, M.; Wegge, P. & Fredriksen, T. (2010). "Do tigers displace leopards? If so, why?". Ecological Research. 25 (4): 875–881. Bibcode:2010EcoR...25..875O. doi:10.1007/s11284-010-0723-1.
Harihar, A.; Pandav, B. & Goyal, S. P. (2011). "Responses of leopard Panthera pardus to the recovery of a tiger Panthera tigris population". Journal of Applied Ecology. 48 (3): 806–814. Bibcode:2011JApEc..48..806H. doi:10.1111/j.1365-2664.2011.01981.x.
Sadleir, R. (1966). "Notes on the Reproduction of the larger Felidae". International Zoo Yearbook. 6: 184–187. doi:10.1111/j.1748-1090.1966.tb01746.x.
Williams, S. T.; Williams, K. S.; Lewis, B. P. & Hill, R. A. (2017). "Population dynamics and threats to an apex predator outside protected areas: implications for carnivore management". Royal Society Open Science. 4 (4) 161090. Bibcode:2017RSOS....461090W. doi:10.1098/rsos.161090. PMC 5414262. PMID 28484625.
Bergin, D. & Nijman, V. (2015). "Potential benefits of impending Moroccan wildlife trade laws, a case study in carnivore skins". Biodiversity and Conservation. 25 (1): 199–201. doi:10.1007/s10531-015-1042-1. S2CID 34533018. Archived from the original on 2020-01-07. Retrieved 2017-01-11.
Äbischer, T.; Ibrahim, T.; Hickisch, R.; Furrer, R. D.; Leuenberger, C. & Wegmann, D. (2020). "Apex predators decline after an influx of pastoralists in former Central African Republic hunting zones" (PDF). Biological Conservation. 241 108326. Bibcode:2020BCons.24108326A. doi:10.1016/j.biocon.2019.108326. S2CID 213766740. Archived (PDF) from the original on 2020-10-03.
Gomez, L. & Shepherd, C.R. (2021). "The illegal exploitation of the Javan Leopard (Panthera pardus melas) and Sunda Clouded Leopard (Neofelis diardi) in Indonesia". Nature Conservation. 43 (43): 25–39. Bibcode:2021NaCon..43...25G. doi:10.3897/natureconservation.43.59399. S2CID 233286106.
Murphey, R. (1951). "The Decline of North Africa Since the Roman Occupation: Climatic or Human?" (PDF). Annals of the Association of American Geographers. XLI (2): 116–132. doi:10.1080/00045605109352048. Archived (PDF) from the original on 2006-09-14.

google.com (Global: 109^th place; English: 87^th place)

Wozencraft, W. C. (2005). "Species Panthera pardus". In Wilson, D. E.; Reeder, D. M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Johns Hopkins University Press. p. 547. ISBN 978-0-8018-8221-0. OCLC 62265494.

hdl.handle.net (Global: 66^th place; English: 34^th place)

Marciszak, A.; Lipecki, G.; Gornig, W.; Matyaszczyk, L.; Oszczepalińska, O.; Nowakowski, D.; Talamo, S. (2022). "The first radiocarbon-dated remains of the Leopard Panthera pardus (Linnaeus, 1758) from the Pleistocene of Poland". Radiocarbon. 64 (6): 1359–1372. Bibcode:2022Radcb..64.1359M. doi:10.1017/RDC.2022.33. hdl:11585/887180. S2CID 255850069. Archived from the original on 2024-05-05. Retrieved 2023-07-18.
Sanchis, A.; Real, C.; Sauqué, V.; Núñez-Lahuerta, C.; Égüez, N.; Tormo, C.; Ripoll, M. P.; Carrión, Y. M.; Duarte, E. & de la Rasilla, M. (2019). "Neanderthal and carnivore activities at Llonin Cave, Asturias, northern Iberian Peninsula: Faunal study of Mousterian levels (MIS 3)". Comptes Rendus Palevol. 18 (1): 113–141. Bibcode:2019CRPal..18..113S. doi:10.1016/j.crpv.2018.06.001. hdl:10651/49880. S2CID 134157124.
Mecozzi, Beniamino; Sardella, Raffaele; Boscaini, Alberto; Cherin, Marco; Costeur, Loïc; Madurell-Malapeira, Joan; Pavia, Marco; Profico, Antonio; Iurino, Dawid A. (15 June 2021). "The tale of a short-tailed cat: New outstanding Late Pleistocene fossils of Lynx pardinus from southern Italy". Quaternary Science Reviews. 262 106840. Bibcode:2021QSRv..26206840M. doi:10.1016/j.quascirev.2021.106840. hdl:2434/959858.
Powell, J.; Axmacher, J. C.; Linnell, J. D.C. & Durant, S. M. (2021). "Diverse locations and a long history: Historical context for urban Leopards (Panthera pardus) in the early anthropocene From Seoul, Korea". Frontiers in Conservation Science. 2 765911. Bibcode:2021FrCS....2.5911P. doi:10.3389/fcosc.2021.765911. hdl:11250/2999764.
Karataş, A.; Bulut, Ş. & Akbaba, B. (2021). "Camera trap records confirm the survival of the Leopard (Panthera pardus L., 1758) in eastern Turkey (Mammalia: Felidae)". Zoology in the Middle East. 67 (3): 198–205. doi:10.1080/09397140.2021.1924419. hdl:11491/7382. S2CID 235564429.
Asad, M.; Waseem, M.; Ross, J. G. & Paterson, A. M. (2019). "The Un-Common Leopard: presence, distribution and abundance in Gallies and Murree Forest Division, Northern Pakistan". Nature Conservation. 37: 53–80. Bibcode:2019NaCon..37...53A. doi:10.3897/natureconservation.37.32748. hdl:10182/11138.
Rostro-García, S.; Kamler, J. F.; Ash, E.; Clements, G. R.; Gibson, L.; Lynam, A. J.; McEwin, R.; Naing, H. & Paglia, S. (2016). "Endangered leopards: Range collapse of the Indochinese leopard (Panthera pardus delacouri) in Southeast Asia". Biological Conservation. 201: 293–300. Bibcode:2016BCons.201..293R. doi:10.1016/j.biocon.2016.07.001. hdl:10722/232870.
Blanco, R. E.; Luzardo-Vera, D. E. (2021). "A biomechanical model to assess the injury risk of leopards (Panthera pardus) hunting by free falling from trees". Biological Communications. 66 (3). doi:10.21638/spbu03.2021.305. hdl:20.500.12008/38512. ISSN 2587-5779.
Rafiq, K.; Wayward, M. W.; Wilson, A. M.; Meloro, C.; Jordan, N. R.; Wich, S. A.; McNutt, J. W.; Golabek, K. A. (2020). "Spatial and temporal overlaps between leopards (Panthera pardus) and their competitors in the African large predator guild". Journal of Zoology. 311 (4): 246–259. doi:10.1111/jzo.12781. hdl:1959.4/unsworks_67550.

ibs.bialowieza.pl (Global: low place; English: low place)

Hayward, M.W.; Henschel, P.; O'Brien, J.; Hofmeyr, M.; Balme, G. & Kerley, G. I. H. (2006). "Prey preferences of the leopard (Panthera pardus)" (PDF). Journal of Zoology. 270 (4): 298–313. doi:10.1111/j.1469-7998.2006.00139.x. Archived (PDF) from the original on 2012-11-05.

indiatimes.com (Global: 31^st place; English: 19^th place)

timesofindia.indiatimes.com

"Is this the longest leopard in India?". The Times of India. 2016. Archived from the original on 2019-04-06. Retrieved 2018-03-14.

iucnredlist.org (Global: 39^th place; English: 71^st place)

Stein, A.B.; Gerngross, P.; Al Hikmani, H.; Balme, G.; Bertola, L.; Drouilly, M.; Farhadinia, M.S.; Feng, L.; Ghoddousi, A.; Henschel, P.; Jhala, Y.; Khorozyan, I.; Kittle, A.; Laguardia, A.; Luo, S.-J.; Mann, G.; Miquelle, D.; Moheb, Z.; Raza, H.; Rostro-García, S.; Shivakumar, S.; Song, D. & Wibisono, H. (2025). "Panthera pardus". IUCN Red List of Threatened Species. 2025 e.T15954A286153337. doi:10.2305/IUCN.UK.2025-2.RLTS.T15954A286153337.en.
Shivakumar, S.; Khettry, A.; Surve, N.; Rahman, H.; Ghimirey, Y.; Tharchen, L.; Zaw, T.; Waseem, M. & Jhala, Y. (2023). "Panthera pardus ssp. fusca". IUCN Red List of Threatened Species. 2023 e.T215195524A215195533. Retrieved 17 March 2024.
Wibisono, H.; Wilianto, E.; Pinondang, I.; Rahman, D.A. & Chandradewi, D. (2021). "Panthera pardus ssp. melas". IUCN Red List of Threatened Species. 2021 e.T15962A50660931. doi:10.2305/IUCN.UK.2021-2.RLTS.T15962A50660931.en. Retrieved 11 November 2023.
Al Hikmani, H.; Spalton, A.; Zafar-ul Islam, M.; al-Johany, A.; Sulayem, M.; Al-Duais, M. & Almalki, A. (2023). "Panthera pardus ssp. nimr". IUCN Red List of Threatened Species. 2023 e.T15958A46767457. Retrieved 17 March 2024.
Ghoddousi, A. & Khorozyan, I. (2023). "Panthera pardus ssp. tulliana". IUCN Red List of Threatened Species. 2023 e.T15961A50660903. doi:10.2305/IUCN.UK.2023-1.RLTS.T15961A50660903.en. Retrieved 17 March 2024.
Rostro-García, S.; Kamler, J.F.; Clements, G.R.; Lynam, A.J. & Naing, H. (2019). "Panthera pardus ssp. delacouri". IUCN Red List of Threatened Species. 2019 e.T124159083A163986056. Retrieved 17 March 2024.
Kittle, A.M. & Watson, A. (2020). "Panthera pardus ssp.kotiya". IUCN Red List of Threatened Species. 2020 e.T15959A50660847. Retrieved 17 March 2024.

jstor.org (Global: 23^rd place; English: 15^th place)

Johnson, K. G.; Wei, W.; Reid, D. G.; Jinchu, H. (1993). "Food habits of Asiatic leopards (Panthera pardus fusca) in Wolong Reserve, Sichuan, China". Journal of Mammalogy. 74 (3): 646–650. Bibcode:1993JMamm..74..646J. doi:10.2307/1382285. JSTOR 1382285.
Karanth, K. U. & Sunquist, M. E. (1995). "Prey selection by tiger, leopard and dhole in tropical forests". Journal of Animal Ecology. 64 (4): 439–450. Bibcode:1995JAnEc..64..439K. doi:10.2307/5647. JSTOR 5647.

monash.edu.au (Global: 9,400^th place; English: 6,694^th place)

biolsci.monash.edu.au

Hunter, L.; Balme, G.; Walker, C.; Pretorius, K. & Rosenberg, K. (2003). "The landscape ecology of leopards (Panthera pardus) in northern KwaZulu-Natal, South Africa: a preliminary project report" (PDF). Ecological Journal. 5: 24–30. Archived from the original (PDF) on March 4, 2009.

nationalgeographic.com (Global: 489^th place; English: 425^th place)

news.nationalgeographic.com

Owen, J. (2005). "Medieval Lion Skulls Reveal Secrets of Tower of London 'Zoo'". National Geographic Magazine. Archived from the original on November 5, 2005. Retrieved 2007-09-05.

nih.gov (Global: 5^th place; English: 5^th place)

pubmed.ncbi.nlm.nih.gov

Uphyrkina, O.; Johnson, E. W.; Quigley, H.; Miquelle, D.; Marker, L.; Bush, M. & O'Brien, S. J. (2001). "Phylogenetics, genome diversity and origin of modern leopard, Panthera pardus" (PDF). Molecular Ecology. 10 (11): 2617–2633. Bibcode:2001MolEc..10.2617U. doi:10.1046/j.0962-1083.2001.01350.x. PMID 11883877. S2CID 304770. Archived (PDF) from the original on 2011-09-10.
Anco, C.; Kolokotronis, S. O.; Henschel, P.; Cunningham, S. W.; Amato, G. & Hekkala, E. (2017). "Historical mitochondrial diversity in African leopards (Panthera pardus) revealed by archival museum specimens". Mitochondrial DNA Part A. 29 (3): 455–473. doi:10.1080/24701394.2017.1307973. PMID 28423965. S2CID 4348541.
Johnson, W. E.; Eizirik, E.; Pecon-Slattery, J.; Murphy, W. J.; Antunes, A.; Teeling, E. & O'Brien, S. J. (2006). "The late Miocene radiation of modern Felidae: a genetic assessment". Science. 311 (5757): 73–77. Bibcode:2006Sci...311...73J. doi:10.1126/science.1122277. PMID 16400146. S2CID 41672825. Archived from the original on 2020-10-04. Retrieved 2019-07-05.
Davis, B. W.; Li, G. & Murphy, W. J. (2010). "Supermatrix and species tree methods resolve phylogenetic relationships within the big cats, Panthera (Carnivora: Felidae)" (PDF). Molecular Phylogenetics and Evolution. 56 (1): 64–76. Bibcode:2010MolPE..56...64D. doi:10.1016/j.ympev.2010.01.036. PMID 20138224. Archived from the original (PDF) on 2016-03-05.
Mazák, J. H.; Christiansen, P.; Kitchener, A. C. & Goswami, A. (2011). "Oldest known pantherine skull and evolution of the tiger". PLOS ONE. 6 (10) e25483. Bibcode:2011PLoSO...625483M. doi:10.1371/journal.pone.0025483. PMC 3189913. PMID 22016768.
Tseng, Z. J.; Wang, X.; Slater, G. J.; Takeuchi, G. T.; Li, Q.; Liu, J. & Xie, G. (2014). "Himalayan fossils of the oldest known pantherine establish ancient origin of big cats". Proceedings of the Royal Society B: Biological Sciences. 281 (1774) 20132686. Bibcode:2014PBioS.28132686T. doi:10.1098/rspb.2013.2686. PMC 3843846. PMID 24225466.
Li, G.; Davis, B. W.; Eizirik, E. & Murphy, W. J. (2016). "Phylogenomic evidence for ancient hybridization in the genomes of living cats (Felidae)". Genome Research. 26 (1): 1–11. doi:10.1101/gr.186668.114. PMC 4691742. PMID 26518481.
Paijmans, J. L. A.; Barlow, A.; Förster, D. W.; Henneberger, K.; Meyer, M.; Nickel, B.; Nagel, D.; Havmøller, R. W.; Baryshnikov, G. F.; Joger, U.; Rosendahl, W. & Hofreiter, M. (2018). "Historical biogeography of the leopard (Panthera pardus) and its extinct Eurasian populations". BMC Evolutionary Biology. 18 (1): 156. Bibcode:2018BMCEE..18..156P. doi:10.1186/s12862-018-1268-0. PMC 6198532. PMID 30348080.
Chi T.-C.; Gan Y.; Yang T.-R. & Chang, C.-H. (2021). "First report of leopard fossils from a limestone cave in Kenting area, southern Taiwan". PeerJ. 9 e12020. doi:10.7717/peerj.12020. PMC 8388558. PMID 34513335.
Sun, X.; Peng, L.; Tsutaya, T.; Jiangzuo, Q.; Hasegawa, Y.; Hou, Y.; Han, Y.; Zhuang, Y.; Ramos Madrigal, J.; Taurozzi, A. J.; Mackie, M.; Trochė, G.; Olsen, J. V.; Cappellini, E.; O'Brien, S. J.; Gilbert, M. T. P.; Yamaguchi, N.; Luo, S.-J. (2026). "The Japanese Archipelago sheltered cave lions, not tigers, during the Late Pleistocene". Proceedings of the National Academy of Sciences of the United States of America. 123 (6) e2523901123. Bibcode:2026PNAS..12323901S. doi:10.1073/pnas.2523901123. PMC 12890994. PMID 41587328.
Allen, W. L.; Cuthill, I. C.; Scott-Samuel, N. E. & Baddeley, R. (2010). "Why the leopard got its spots: relating pattern development to ecology in felids". Proceedings of the Royal Society B. 278 (1710): 1373–1380. doi:10.1098/rspb.2010.1734. PMC 3061134. PMID 20961899.
Robinson, R. (1970). "Inheritance of the black form of the leopard Panthera pardus". Genetica. 41 (1): 190–197. doi:10.1007/BF00958904. PMID 5480762. S2CID 5446868.
Eizirik, E.; Yuhki, N.; Johnson, W. E.; Menotti-Raymond, M.; Hannah, S. S.; O'Brien, S. J. (2003). "Molecular genetics and evolution of melanism in the cat family". Current Biology. 13 (5): 448–453. Bibcode:2003CBio...13..448E. doi:10.1016/S0960-9822(03)00128-3. PMID 12620197. S2CID 19021807.
Da Silva L. G., K.; Kawanishi, K.; Henschel P.; Kittle, A.; Sanei, A.; Reebin, A.; Miquelle, D.; Stein, A. B.; Watson, A.; Kekule, L. B.; Machado, R. B. & Eizirik, E. (2017). "Mapping black panthers: Macroecological modeling of melanism in leopards (Panthera pardus)". PLOS ONE. 12 (4) e0170378. Bibcode:2017PLoSO..1270378D. doi:10.1371/journal.pone.0170378. PMC 5381760. PMID 28379961.
Jacobson, A. P.; Gerngross, P.; Lemeris, J. R. Jr.; Schoonover, R. F.; Anco, C.; Breitenmoser-Würsten, C.; Durant, S. M.; Farhadinia, M. S.; Henschel, P.; Kamler, J. F.; Laguardia, A.; Rostro-García, S.; Stein, A. B. & Dollar, L. (2016). "Leopard (Panthera pardus) status, distribution, and the research efforts across its range". PeerJ. 4 e1974. doi:10.7717/peerj.1974. PMC 4861552. PMID 27168983.
Pirie, T. J.; Thomas, R. L. & Fellowes, M. D. E. (2017). "Increasing game prices may alter farmers' behaviours towards leopards (Panthera pardus) and other carnivores in South Africa". PeerJ. 5 e3369. doi:10.7717/peerj.3369. PMC 5452990. PMID 28584709.
Wibisono, H. T.; Wahyudi, H. A.; Wilianto, E.; Pinondang, I. M. R.; Primajati, M.; Liswanto, D. & Linkie, M. (2018). "Identifying priority conservation landscapes and actions for the Critically Endangered Javan leopard in Indonesia: Conserving the last large carnivore in Java Island". PLOS ONE. 13 (6) e0198369. Bibcode:2018PLoSO..1398369W. doi:10.1371/journal.pone.0198369. PMC 6021038. PMID 29949588.
Sauqué, Víctor; Rabal-Garcés, Raquel; Sola-Almagro, Cristina; Cuenca-Bescós, Gloria (18 March 2014). Rook, Lorenzo (ed.). "Bone Accumulation by Leopards in the Late Pleistocene in the Moncayo Massif (Zaragoza, NE Spain)". PLoS ONE. 9 (3) e92144. Bibcode:2014PLoSO...992144S. doi:10.1371/journal.pone.0092144. ISSN 1932-6203. PMC 3958443. PMID 24642667.
Balme, G. A.; Miller, J. R. B.; Pitman, R. T.; Hunter, L. T. B. (2017). "Caching reduces kleptoparasitism in a solitary, large felid". Journal of Animal Ecology. 86 (3): 634–644. Bibcode:2017JAnEc..86..634B. doi:10.1111/1365-2656.12654. PMID 28217865.
Vissia, S.; Fattebert, J.; van Langevelde, F. (2022). "Leopard density and interspecific spatiotemporal interactions in a hyena-dominated landscape". Ecology and Evolution. 12 (10) e9365. Bibcode:2022EcoEv..12E9365V. doi:10.1002/ece3.9365. PMC 9534747. PMID 36225822.
Miller, J. R. B.; Pitman, R. T.; Mann, G. K. H.; Fuller, A. K.; Balme, G. A. (2018). "Lions and leopards coexist without spatial, temporal or demographic effects of interspecific competition". Journal of Animal Ecology. 87 (6): 1709–1726. Bibcode:2018JAnEc..87.1709M. doi:10.1111/1365-2656.12883. PMID 30010193.
Srivathsa, A.; Ramachandran, V.; Saravanan, P.; Sureshbabu, A.; Ganguly, D. & Ramakrishnan, U. (2023). "Topcats and underdogs: intraguild interactions among three apex carnivores across Asia's forestscapes". Biological Reviews. 98 (6): 2114–2135. doi:10.1111/brv.12998. PMID 37449566. S2CID 259903849.
Steinmetz, R.; Seuaturien, N.; Intanajitjuy, P.; Inrueang, P. & Prempree, K. (2021). "The effects of prey depletion on dietary niches of sympatric apex predators in Southeast Asia". Integrative Zoology. 16 (1): 19–32. doi:10.1111/1749-4877.12461. PMID 32627329.
Williams, S. T.; Williams, K. S.; Lewis, B. P. & Hill, R. A. (2017). "Population dynamics and threats to an apex predator outside protected areas: implications for carnivore management". Royal Society Open Science. 4 (4) 161090. Bibcode:2017RSOS....461090W. doi:10.1098/rsos.161090. PMC 5414262. PMID 28484625.

ncbi.nlm.nih.gov

Mazák, J. H.; Christiansen, P.; Kitchener, A. C. & Goswami, A. (2011). "Oldest known pantherine skull and evolution of the tiger". PLOS ONE. 6 (10) e25483. Bibcode:2011PLoSO...625483M. doi:10.1371/journal.pone.0025483. PMC 3189913. PMID 22016768.
Tseng, Z. J.; Wang, X.; Slater, G. J.; Takeuchi, G. T.; Li, Q.; Liu, J. & Xie, G. (2014). "Himalayan fossils of the oldest known pantherine establish ancient origin of big cats". Proceedings of the Royal Society B: Biological Sciences. 281 (1774) 20132686. Bibcode:2014PBioS.28132686T. doi:10.1098/rspb.2013.2686. PMC 3843846. PMID 24225466.
Li, G.; Davis, B. W.; Eizirik, E. & Murphy, W. J. (2016). "Phylogenomic evidence for ancient hybridization in the genomes of living cats (Felidae)". Genome Research. 26 (1): 1–11. doi:10.1101/gr.186668.114. PMC 4691742. PMID 26518481.
Paijmans, J. L. A.; Barlow, A.; Förster, D. W.; Henneberger, K.; Meyer, M.; Nickel, B.; Nagel, D.; Havmøller, R. W.; Baryshnikov, G. F.; Joger, U.; Rosendahl, W. & Hofreiter, M. (2018). "Historical biogeography of the leopard (Panthera pardus) and its extinct Eurasian populations". BMC Evolutionary Biology. 18 (1): 156. Bibcode:2018BMCEE..18..156P. doi:10.1186/s12862-018-1268-0. PMC 6198532. PMID 30348080.
Chi T.-C.; Gan Y.; Yang T.-R. & Chang, C.-H. (2021). "First report of leopard fossils from a limestone cave in Kenting area, southern Taiwan". PeerJ. 9 e12020. doi:10.7717/peerj.12020. PMC 8388558. PMID 34513335.
Sun, X.; Peng, L.; Tsutaya, T.; Jiangzuo, Q.; Hasegawa, Y.; Hou, Y.; Han, Y.; Zhuang, Y.; Ramos Madrigal, J.; Taurozzi, A. J.; Mackie, M.; Trochė, G.; Olsen, J. V.; Cappellini, E.; O'Brien, S. J.; Gilbert, M. T. P.; Yamaguchi, N.; Luo, S.-J. (2026). "The Japanese Archipelago sheltered cave lions, not tigers, during the Late Pleistocene". Proceedings of the National Academy of Sciences of the United States of America. 123 (6) e2523901123. Bibcode:2026PNAS..12323901S. doi:10.1073/pnas.2523901123. PMC 12890994. PMID 41587328.
Allen, W. L.; Cuthill, I. C.; Scott-Samuel, N. E. & Baddeley, R. (2010). "Why the leopard got its spots: relating pattern development to ecology in felids". Proceedings of the Royal Society B. 278 (1710): 1373–1380. doi:10.1098/rspb.2010.1734. PMC 3061134. PMID 20961899.
Stein, A. B. & Hayssen, V. (2010). "Panthera pardus (Carnivora: Felidae)". Mammalian Species. 45 (900): 30–48. doi:10.1644/900.1. PMC 7149468. S2CID 44839740.
Da Silva L. G., K.; Kawanishi, K.; Henschel P.; Kittle, A.; Sanei, A.; Reebin, A.; Miquelle, D.; Stein, A. B.; Watson, A.; Kekule, L. B.; Machado, R. B. & Eizirik, E. (2017). "Mapping black panthers: Macroecological modeling of melanism in leopards (Panthera pardus)". PLOS ONE. 12 (4) e0170378. Bibcode:2017PLoSO..1270378D. doi:10.1371/journal.pone.0170378. PMC 5381760. PMID 28379961.
Jacobson, A. P.; Gerngross, P.; Lemeris, J. R. Jr.; Schoonover, R. F.; Anco, C.; Breitenmoser-Würsten, C.; Durant, S. M.; Farhadinia, M. S.; Henschel, P.; Kamler, J. F.; Laguardia, A.; Rostro-García, S.; Stein, A. B. & Dollar, L. (2016). "Leopard (Panthera pardus) status, distribution, and the research efforts across its range". PeerJ. 4 e1974. doi:10.7717/peerj.1974. PMC 4861552. PMID 27168983.
Pirie, T. J.; Thomas, R. L. & Fellowes, M. D. E. (2017). "Increasing game prices may alter farmers' behaviours towards leopards (Panthera pardus) and other carnivores in South Africa". PeerJ. 5 e3369. doi:10.7717/peerj.3369. PMC 5452990. PMID 28584709.
Wibisono, H. T.; Wahyudi, H. A.; Wilianto, E.; Pinondang, I. M. R.; Primajati, M.; Liswanto, D. & Linkie, M. (2018). "Identifying priority conservation landscapes and actions for the Critically Endangered Javan leopard in Indonesia: Conserving the last large carnivore in Java Island". PLOS ONE. 13 (6) e0198369. Bibcode:2018PLoSO..1398369W. doi:10.1371/journal.pone.0198369. PMC 6021038. PMID 29949588.
Sauqué, Víctor; Rabal-Garcés, Raquel; Sola-Almagro, Cristina; Cuenca-Bescós, Gloria (18 March 2014). Rook, Lorenzo (ed.). "Bone Accumulation by Leopards in the Late Pleistocene in the Moncayo Massif (Zaragoza, NE Spain)". PLoS ONE. 9 (3) e92144. Bibcode:2014PLoSO...992144S. doi:10.1371/journal.pone.0092144. ISSN 1932-6203. PMC 3958443. PMID 24642667.
Vissia, S.; Fattebert, J.; van Langevelde, F. (2022). "Leopard density and interspecific spatiotemporal interactions in a hyena-dominated landscape". Ecology and Evolution. 12 (10) e9365. Bibcode:2022EcoEv..12E9365V. doi:10.1002/ece3.9365. PMC 9534747. PMID 36225822.
Williams, S. T.; Williams, K. S.; Lewis, B. P. & Hill, R. A. (2017). "Population dynamics and threats to an apex predator outside protected areas: implications for carnivore management". Royal Society Open Science. 4 (4) 161090. Bibcode:2017RSOS....461090W. doi:10.1098/rsos.161090. PMC 5414262. PMID 28484625.

nus.edu.sg (Global: 2,453^rd place; English: 1,629^th place)

science.nus.edu.sg

Gray, T. N. & Phan, C. (2011). "Habitat preferences and activity patterns of the larger mammal community in Phnom Prich Wildlife Sanctuary, Cambodia" (PDF). The Raffles Bulletin of Zoology. 59 (2): 311−318. Archived from the original on 2024-05-05. Retrieved 2019-04-16.

openedition.org (Global: 366^th place; English: 883^rd place)

journals.openedition.org

Sauqué, V. & Cuenca-Bescós, G. (2013). "The Iberian Peninsula, the last European refugium of Panthera pardus Linnaeus 1758 during the Upper Pleistocene". Quaternaire. 24 (1): 13–24. doi:10.4000/quaternaire.6468. Archived from the original on 2023-07-18. Retrieved 2023-07-18.

palmbeachpost.com (Global: 3,040^th place; English: 1,633^rd place)

Salisbury, S. (2014). "Roxanne, oldest spotted leopard in captivity, dies at Acreage preserve". The Palm Beach Post. Archived from the original on 2014-08-11.

rero.ch (Global: 3,023^rd place; English: 2,092^nd place)

doc.rero.ch

Äbischer, T.; Ibrahim, T.; Hickisch, R.; Furrer, R. D.; Leuenberger, C. & Wegmann, D. (2020). "Apex predators decline after an influx of pastoralists in former Central African Republic hunting zones" (PDF). Biological Conservation. 241 108326. Bibcode:2020BCons.24108326A. doi:10.1016/j.biocon.2019.108326. S2CID 213766740. Archived (PDF) from the original on 2020-10-03.

researchgate.net (Global: 96^th place; English: 89^th place)

Werdelin, L.; Yamaguchi, N.; Johnson, W. E. & O'Brien, S. J. (2010). "Phylogeny and evolution of cats (Felidae)". In Macdonald, D. W. & Loveridge, A. J. (eds.). Biology and Conservation of Wild Felids. Oxford, UK: Oxford University Press. pp. 59–82. ISBN 978-0-19-923445-5. Archived from the original on 2018-09-25. Retrieved 2018-09-25.
Marciszak, A. & Stefaniak, K. (2010). "Two forms of cave lion: Middle Pleistocene Panthera spelaea fossilis Reichenau, 1906 and Upper Pleistocene Panthera spelaea spelaea Goldfuss, 1810 from the Bísnik Cave, Poland". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 258 (3): 339–351. Bibcode:2010NJGPA.258..339M. doi:10.1127/0077-7749/2010/0117. Archived from the original on 2018-09-25. Retrieved 2017-08-24.
Jenny, D. & Zuberbuhler, K. (2005). "Hunting behaviour in west African forest leopards". African Journal of Ecology. 43 (3): 197–200. Bibcode:2005AfJEc..43..197J. doi:10.1111/j.1365-2028.2005.00565.x. Archived from the original on 2018-09-25. Retrieved 2018-09-25.
Jenny, D. (1996). "Spatial organization of leopards Panthera pardus in Tai National Park, Ivory Coast: Is rainforest habitat a "tropical haven"?". Journal of Zoology. 240 (3): 427–440. Bibcode:1996JZoo..240..427J. doi:10.1111/j.1469-7998.1996.tb05296.x. Archived from the original on 2024-05-05. Retrieved 2021-09-06.
Blanco, R. E.; Luzardo-Vera, D. E. (2021). "A biomechanical model to assess the injury risk of leopards (Panthera pardus) hunting by free falling from trees". Biological Communications. 66 (3). doi:10.21638/spbu03.2021.305. hdl:20.500.12008/38512. ISSN 2587-5779.
Bergin, D. & Nijman, V. (2014). "Open, Unregulated Trade in Wildlife in Morocco's Markets". TRAFFIC Bulletin. 26 (1): 65–70. Archived from the original on 2018-10-31. Retrieved 2015-03-23.
Bergin, D. & Nijman, V. (2015). "Potential benefits of impending Moroccan wildlife trade laws, a case study in carnivore skins". Biodiversity and Conservation. 25 (1): 199–201. doi:10.1007/s10531-015-1042-1. S2CID 34533018. Archived from the original on 2020-01-07. Retrieved 2017-01-11.

sabinet.co.za (Global: low place; English: low place)

reference.sabinet.co.za

Marker, L. L. & Dickman, A. J. (2005). "Factors affecting leopard (Panthera pardus) spatial ecology, with particular reference to Namibian farmlands" (PDF). South African Journal of Wildlife Research. 35 (2): 105–115.
Bothma, J. du P. (2005). "Water-use by southern Kalahari leopards" (PDF). South African Journal of Wildlife Research. 35: 131–137.

semanticscholar.org (Global: 15^th place; English: 8^th place)

api.semanticscholar.org

Nicholas, N. (1999). "A conundrum of cats: pards and their relatives in Byzantium". Greek, Roman, and Byzantine Studies. 40: 253–298. S2CID 56160515.
Uphyrkina, O.; Johnson, E. W.; Quigley, H.; Miquelle, D.; Marker, L.; Bush, M. & O'Brien, S. J. (2001). "Phylogenetics, genome diversity and origin of modern leopard, Panthera pardus" (PDF). Molecular Ecology. 10 (11): 2617–2633. Bibcode:2001MolEc..10.2617U. doi:10.1046/j.0962-1083.2001.01350.x. PMID 11883877. S2CID 304770. Archived (PDF) from the original on 2011-09-10.
Anco, C.; Kolokotronis, S. O.; Henschel, P.; Cunningham, S. W.; Amato, G. & Hekkala, E. (2017). "Historical mitochondrial diversity in African leopards (Panthera pardus) revealed by archival museum specimens". Mitochondrial DNA Part A. 29 (3): 455–473. doi:10.1080/24701394.2017.1307973. PMID 28423965. S2CID 4348541.
Johnson, W. E.; Eizirik, E.; Pecon-Slattery, J.; Murphy, W. J.; Antunes, A.; Teeling, E. & O'Brien, S. J. (2006). "The late Miocene radiation of modern Felidae: a genetic assessment". Science. 311 (5757): 73–77. Bibcode:2006Sci...311...73J. doi:10.1126/science.1122277. PMID 16400146. S2CID 41672825. Archived from the original on 2020-10-04. Retrieved 2019-07-05.
Marciszak, A.; Lipecki, G.; Gornig, W.; Matyaszczyk, L.; Oszczepalińska, O.; Nowakowski, D.; Talamo, S. (2022). "The first radiocarbon-dated remains of the Leopard Panthera pardus (Linnaeus, 1758) from the Pleistocene of Poland". Radiocarbon. 64 (6): 1359–1372. Bibcode:2022Radcb..64.1359M. doi:10.1017/RDC.2022.33. hdl:11585/887180. S2CID 255850069. Archived from the original on 2024-05-05. Retrieved 2023-07-18.
Sanchis, A.; Real, C.; Sauqué, V.; Núñez-Lahuerta, C.; Égüez, N.; Tormo, C.; Ripoll, M. P.; Carrión, Y. M.; Duarte, E. & de la Rasilla, M. (2019). "Neanderthal and carnivore activities at Llonin Cave, Asturias, northern Iberian Peninsula: Faunal study of Mousterian levels (MIS 3)". Comptes Rendus Palevol. 18 (1): 113–141. Bibcode:2019CRPal..18..113S. doi:10.1016/j.crpv.2018.06.001. hdl:10651/49880. S2CID 134157124.
Stein, A. B. & Hayssen, V. (2010). "Panthera pardus (Carnivora: Felidae)". Mammalian Species. 45 (900): 30–48. doi:10.1644/900.1. PMC 7149468. S2CID 44839740.
Robinson, R. (1970). "Inheritance of the black form of the leopard Panthera pardus". Genetica. 41 (1): 190–197. doi:10.1007/BF00958904. PMID 5480762. S2CID 5446868.
Eizirik, E.; Yuhki, N.; Johnson, W. E.; Menotti-Raymond, M.; Hannah, S. S.; O'Brien, S. J. (2003). "Molecular genetics and evolution of melanism in the cat family". Current Biology. 13 (5): 448–453. Bibcode:2003CBio...13..448E. doi:10.1016/S0960-9822(03)00128-3. PMID 12620197. S2CID 19021807.
Soultan, A.; Attum, O.; Hamada, A.; Hatab, E. B.; Ahmed, S. E.; Eisa, A.; Al Sharif, I.; Nagy, A. & Shohdi, W. (2017). "Recent observation for leopard Panthera pardus in Egypt". Mammalia. 81 (1): 115–117. doi:10.1515/mammalia-2015-0089. S2CID 90676105.
Karataş, A.; Bulut, Ş. & Akbaba, B. (2021). "Camera trap records confirm the survival of the Leopard (Panthera pardus L., 1758) in eastern Turkey (Mammalia: Felidae)". Zoology in the Middle East. 67 (3): 198–205. doi:10.1080/09397140.2021.1924419. hdl:11491/7382. S2CID 235564429.
Johnson, A.; Vongkhamheng, C.; Hedemark, M. & Saithongdam, T. (2006). "Effects of human–carnivore conflict on tiger (Panthera tigris) and prey populations in Lao PDR" (PDF). Animal Conservation. 9 (4): 421–430. Bibcode:2006AnCon...9..421J. doi:10.1111/j.1469-1795.2006.00049.x. S2CID 73637721. Archived (PDF) from the original on 2017-08-10.
Odden, M. & Wegge, P. (2005). "Spacing and activity patterns of leopards Panthera pardus in the Royal Bardia National Park, Nepal". Wildlife Biology. 11 (2): 145–152. doi:10.2981/0909-6396(2005)11[145:SAAPOL]2.0.CO;2. S2CID 86140708.
Kruuk, H. & Turner, M. (1967). "Comparative notes on predation by lion, leopard, cheetah and wild dog in the Serengeti area, East Africa". Mammalia. 31 (1): 1–27. doi:10.1515/mamm.1967.31.1.1. S2CID 84619500.
Srivathsa, A.; Ramachandran, V.; Saravanan, P.; Sureshbabu, A.; Ganguly, D. & Ramakrishnan, U. (2023). "Topcats and underdogs: intraguild interactions among three apex carnivores across Asia's forestscapes". Biological Reviews. 98 (6): 2114–2135. doi:10.1111/brv.12998. PMID 37449566. S2CID 259903849.
Bergin, D. & Nijman, V. (2015). "Potential benefits of impending Moroccan wildlife trade laws, a case study in carnivore skins". Biodiversity and Conservation. 25 (1): 199–201. doi:10.1007/s10531-015-1042-1. S2CID 34533018. Archived from the original on 2020-01-07. Retrieved 2017-01-11.
Äbischer, T.; Ibrahim, T.; Hickisch, R.; Furrer, R. D.; Leuenberger, C. & Wegmann, D. (2020). "Apex predators decline after an influx of pastoralists in former Central African Republic hunting zones" (PDF). Biological Conservation. 241 108326. Bibcode:2020BCons.24108326A. doi:10.1016/j.biocon.2019.108326. S2CID 213766740. Archived (PDF) from the original on 2020-10-03.
Gomez, L. & Shepherd, C.R. (2021). "The illegal exploitation of the Javan Leopard (Panthera pardus melas) and Sunda Clouded Leopard (Neofelis diardi) in Indonesia". Nature Conservation. 43 (43): 25–39. Bibcode:2021NaCon..43...25G. doi:10.3897/natureconservation.43.59399. S2CID 233286106.

si.edu (Global: 256^th place; English: 197^th place)

repository.si.edu

Miththapala, S.; Seidensticker, J. & O'Brien, S. J. (1996). "Phylogeographic subspecies recognition in leopards (Panthera pardus): molecular genetic variation" (PDF). Conservation Biology. 10 (4): 1115–1132. Bibcode:1996ConBi..10.1115M. doi:10.1046/j.1523-1739.1996.10041115.x. Archived (PDF) from the original on 2016-02-17. Retrieved 2017-03-09.
Kitchener, A. C.; Breitenmoser-Würsten, C.; Eizirik, E.; Gentry, A.; Werdelin, L.; Wilting, A.; Yamaguchi, N.; Abramov, A. V.; Christiansen, P.; Driscoll, C.; Duckworth, J. W.; Johnson, W.; Luo, S.-J.; Meijaard, E.; O'Donoghue, P.; Sanderson, J.; Seymour, K.; Bruford, M.; Groves, C.; Hoffmann, M.; Nowell, K.; Timmons, Z. & Tobe, S. (2017). "A revised taxonomy of the Felidae: The final report of the Cat Classification Task Force of the IUCN Cat Specialist Group" (PDF). Cat News (Special Issue 11): 73–75. Archived (PDF) from the original on 2020-01-17. Retrieved 2019-07-25.

siamese-heritage.org (Global: low place; English: low place)

Gray, T. N. E. (2013). "Activity patterns and home ranges of Indochinese leopard Panthera pardus delacouri in the Eastern Plains Landscape, Cambodia" (PDF). Natural History Bulletin of the Siam Society. 59: 39−47. Archived (PDF) from the original on 2016-02-22.

studylib.net (Global: low place; English: low place)

Robichaud, W.; Insua-Cao; Sisomphane, P. C. & Chounnavanh, S. (2010). "Appendix 4". A scoping mission to Nam Kan National Protected Area, Lao PDR. Fauna & Flora International. pp. 33−42. Archived from the original on 2019-04-07. Retrieved 2019-04-07.

theshortstory.co.uk (Global: low place; English: low place)

Kipling, R. (1902). "How the Leopard Got His Spots" (PDF). Just So Stories. Macmillan. Archived (PDF) from the original on 2023-11-29. Retrieved 2023-05-14.

traffic.org (Global: low place; English: low place)

Raza, R.H.; Chauhan, D.S.; Pasha, M.K.S. & Sinha, S. (2012). Illuminating the blind spot: A study on illegal trade in Leopard parts in India (2001–2010) (PDF) (Report). New Delhi: TRAFFIC India, WWF India. Archived (PDF) from the original on 2020-09-24.

uio.no (Global: 2,380^th place; English: 2,625^th place)

lynx.uio.no

Nowell, K. & Jackson, P. (1996). "Leopard Panthera pardus (Linnaeus, 1758)". Wild Cats: status survey and conservation action plan. Gland, Switzerland: IUCN/SSC Cat Specialist Group. Archived from the original on 2014-02-22.

web.archive.org (Global: 1^st place; English: 1^st place)

Oken, L. (1816). "1. Art, Panthera". Lehrbuch der Zoologie. 2. Abtheilung. Jena: August Schmid & Comp. p. 1052. Archived from the original on 2024-05-05. Retrieved 2019-04-06.
Allen, J. A. (1902). "Mammal names proposed by Oken in his 'Lehrbuch der Zoologie'". Bulletin of the American Museum of Natural History. 16 (27): 373−379. Archived from the original on 2023-12-09. Retrieved 2024-03-14.
Miththapala, S.; Seidensticker, J. & O'Brien, S. J. (1996). "Phylogeographic subspecies recognition in leopards (Panthera pardus): molecular genetic variation" (PDF). Conservation Biology. 10 (4): 1115–1132. Bibcode:1996ConBi..10.1115M. doi:10.1046/j.1523-1739.1996.10041115.x. Archived (PDF) from the original on 2016-02-17. Retrieved 2017-03-09.
Uphyrkina, O.; Johnson, E. W.; Quigley, H.; Miquelle, D.; Marker, L.; Bush, M. & O'Brien, S. J. (2001). "Phylogenetics, genome diversity and origin of modern leopard, Panthera pardus" (PDF). Molecular Ecology. 10 (11): 2617–2633. Bibcode:2001MolEc..10.2617U. doi:10.1046/j.0962-1083.2001.01350.x. PMID 11883877. S2CID 304770. Archived (PDF) from the original on 2011-09-10.
Kitchener, A. C.; Breitenmoser-Würsten, C.; Eizirik, E.; Gentry, A.; Werdelin, L.; Wilting, A.; Yamaguchi, N.; Abramov, A. V.; Christiansen, P.; Driscoll, C.; Duckworth, J. W.; Johnson, W.; Luo, S.-J.; Meijaard, E.; O'Donoghue, P.; Sanderson, J.; Seymour, K.; Bruford, M.; Groves, C.; Hoffmann, M.; Nowell, K.; Timmons, Z. & Tobe, S. (2017). "A revised taxonomy of the Felidae: The final report of the Cat Classification Task Force of the IUCN Cat Specialist Group" (PDF). Cat News (Special Issue 11): 73–75. Archived (PDF) from the original on 2020-01-17. Retrieved 2019-07-25.
Meyer, F. A. A. (1794). "Über de la Metheries schwarzen Panther". Zoologische Annalen. Erster Band. Weimar: Im Verlage des Industrie-Comptoirs. pp. 394–396. Archived from the original on 2024-05-05. Retrieved 2019-04-05.
Spalton, J. A. & Al Hikmani, H. M. (2006). "The Leopard in the Arabian Peninsula – Distribution and subspecies status" (PDF). Cat News (Special Issue 1): 4–8. Archived (PDF) from the original on 2015-06-19.
Johnson, W. E.; Eizirik, E.; Pecon-Slattery, J.; Murphy, W. J.; Antunes, A.; Teeling, E. & O'Brien, S. J. (2006). "The late Miocene radiation of modern Felidae: a genetic assessment". Science. 311 (5757): 73–77. Bibcode:2006Sci...311...73J. doi:10.1126/science.1122277. PMID 16400146. S2CID 41672825. Archived from the original on 2020-10-04. Retrieved 2019-07-05.
Werdelin, L.; Yamaguchi, N.; Johnson, W. E. & O'Brien, S. J. (2010). "Phylogeny and evolution of cats (Felidae)". In Macdonald, D. W. & Loveridge, A. J. (eds.). Biology and Conservation of Wild Felids. Oxford, UK: Oxford University Press. pp. 59–82. ISBN 978-0-19-923445-5. Archived from the original on 2018-09-25. Retrieved 2018-09-25.
Davis, B. W.; Li, G. & Murphy, W. J. (2010). "Supermatrix and species tree methods resolve phylogenetic relationships within the big cats, Panthera (Carnivora: Felidae)" (PDF). Molecular Phylogenetics and Evolution. 56 (1): 64–76. Bibcode:2010MolPE..56...64D. doi:10.1016/j.ympev.2010.01.036. PMID 20138224. Archived from the original (PDF) on 2016-03-05.
Marciszak, A.; Lipecki, G.; Gornig, W.; Matyaszczyk, L.; Oszczepalińska, O.; Nowakowski, D.; Talamo, S. (2022). "The first radiocarbon-dated remains of the Leopard Panthera pardus (Linnaeus, 1758) from the Pleistocene of Poland". Radiocarbon. 64 (6): 1359–1372. Bibcode:2022Radcb..64.1359M. doi:10.1017/RDC.2022.33. hdl:11585/887180. S2CID 255850069. Archived from the original on 2024-05-05. Retrieved 2023-07-18.
Sauqué, V. & Cuenca-Bescós, G. (2013). "The Iberian Peninsula, the last European refugium of Panthera pardus Linnaeus 1758 during the Upper Pleistocene". Quaternaire. 24 (1): 13–24. doi:10.4000/quaternaire.6468. Archived from the original on 2023-07-18. Retrieved 2023-07-18.
Marciszak, A. & Stefaniak, K. (2010). "Two forms of cave lion: Middle Pleistocene Panthera spelaea fossilis Reichenau, 1906 and Upper Pleistocene Panthera spelaea spelaea Goldfuss, 1810 from the Bísnik Cave, Poland". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 258 (3): 339–351. Bibcode:2010NJGPA.258..339M. doi:10.1127/0077-7749/2010/0117. Archived from the original on 2018-09-25. Retrieved 2017-08-24.
Kawata, K. (2001). "Zoological gardens of Japan". In Kisling, V.N. (ed.). Zoo and Aquarium History: Ancient Animal Collections to Zoological Gardens. Boca Raton, Florida: CRC Press. pp. 295–329. ISBN 978-0-8493-2100-9. Archived from the original on 2024-05-05. Retrieved 2021-09-07.
Mills, M. G. L. (2005). "Subfamily Pantherinae". In Skinner, J. D.; Chimimba, C. T. (eds.). The mammals of the southern African sub region (Third ed.). Cambridge: Cambridge University Press. pp. 385–396. ISBN 978-0-521-84418-5. Archived from the original on 2024-05-05. Retrieved 2021-05-19.
Nowell, K. & Jackson, P. (1996). "Leopard Panthera pardus (Linnaeus, 1758)". Wild Cats: status survey and conservation action plan. Gland, Switzerland: IUCN/SSC Cat Specialist Group. Archived from the original on 2014-02-22.
Hoath, R. (2009). "Leopard Panthera pardus (Linnaeus, 1758)". Field Guide to the Mammals of Egypt. Cairo, Egypt: American University in Cairo Press. pp. 106–107. ISBN 978-977-416-254-1. Archived from the original on 2024-05-05. Retrieved 2021-05-19.
Estes, R. (1991). "Leopard Panthera pardus". The Behavior Guide to African Mammals, Including Hoofed Mammals, Carnivores, Primates. Los Angeles: The University of California Press. pp. 366–369. ISBN 978-0-520-08085-0. Archived from the original on 2024-05-05. Retrieved 2021-05-19.
Hunter, L.; Henschel, P. & Ray, J. C. (2013). "Panthera pardus Leopard". In Kingdon, J.; Happold, D.; Butynski, T.; Hoffmann, M.; Happold, M. & Kalina, J. (eds.). Mammals of Africa. London: Bloomsbury Publishing. pp. 159–168. ISBN 978-1-4081-8996-2. Archived from the original on 2024-05-05. Retrieved 2021-05-19.
Nowak, R. M. (1999). "Panthera pardus (Leopard)". Walker's Mammals of the World (Sixth ed.). Baltimore, US: Johns Hopkins University Press. pp. 828–831. ISBN 978-0-8018-5789-8. Archived from the original on 2024-05-05. Retrieved 2021-05-19.
"Is this the longest leopard in India?". The Times of India. 2016. Archived from the original on 2019-04-06. Retrieved 2018-03-14.
Prater, S. H. (1921). "Record panther skull (P. p. pardus)". The Journal of the Bombay Natural History Society. XXVII (Part IV): 933–935. Archived from the original on 2019-05-02. Retrieved 2019-08-24.
Judas, J.; Paillat, P.; Khoja, A. & Boug, A. (2006). "Status of the Arabian leopard in Saudi Arabia" (PDF). Cat News (Special Issue 1): 11–19. Archived from the original (PDF) on 2015-09-19. Retrieved 2019-03-17.
Arthreya, V. (2012). "Living with Leopards Outside Protected Areas in India". Conservation India. Archived from the original on 2012-12-21. Retrieved 2012-12-10.
Johnson, A.; Vongkhamheng, C.; Hedemark, M. & Saithongdam, T. (2006). "Effects of human–carnivore conflict on tiger (Panthera tigris) and prey populations in Lao PDR" (PDF). Animal Conservation. 9 (4): 421–430. Bibcode:2006AnCon...9..421J. doi:10.1111/j.1469-1795.2006.00049.x. S2CID 73637721. Archived (PDF) from the original on 2017-08-10.
Robichaud, W.; Insua-Cao; Sisomphane, P. C. & Chounnavanh, S. (2010). "Appendix 4". A scoping mission to Nam Kan National Protected Area, Lao PDR. Fauna & Flora International. pp. 33−42. Archived from the original on 2019-04-07. Retrieved 2019-04-07.
Gray, T. N. & Phan, C. (2011). "Habitat preferences and activity patterns of the larger mammal community in Phnom Prich Wildlife Sanctuary, Cambodia" (PDF). The Raffles Bulletin of Zoology. 59 (2): 311−318. Archived from the original on 2024-05-05. Retrieved 2019-04-16.
Gray, T. N. E. (2013). "Activity patterns and home ranges of Indochinese leopard Panthera pardus delacouri in the Eastern Plains Landscape, Cambodia" (PDF). Natural History Bulletin of the Siam Society. 59: 39−47. Archived (PDF) from the original on 2016-02-22.
Hunter, L.; Balme, G.; Walker, C.; Pretorius, K. & Rosenberg, K. (2003). "The landscape ecology of leopards (Panthera pardus) in northern KwaZulu-Natal, South Africa: a preliminary project report" (PDF). Ecological Journal. 5: 24–30. Archived from the original (PDF) on March 4, 2009.
Jenny, D. & Zuberbuhler, K. (2005). "Hunting behaviour in west African forest leopards". African Journal of Ecology. 43 (3): 197–200. Bibcode:2005AfJEc..43..197J. doi:10.1111/j.1365-2028.2005.00565.x. Archived from the original on 2018-09-25. Retrieved 2018-09-25.
Sunquist, M. E. & Sunquist, F. (2002). "Leopard Panthera pardus". Wild Cats of the World. Chicago: University of Chicago Press. pp. 318–342. ISBN 978-0-226-77999-7. Archived from the original on 2024-04-30. Retrieved 2021-05-19.
Bothma, Jacobus du P.; Walker, Clive (2013-11-11). Larger Carnivores of the African Savannas. Springer Science & Business Media. ISBN 978-3-662-03766-9. Archived from the original on 2024-01-06. Retrieved 2024-01-06.
Jenny, D. (1996). "Spatial organization of leopards Panthera pardus in Tai National Park, Ivory Coast: Is rainforest habitat a "tropical haven"?". Journal of Zoology. 240 (3): 427–440. Bibcode:1996JZoo..240..427J. doi:10.1111/j.1469-7998.1996.tb05296.x. Archived from the original on 2024-05-05. Retrieved 2021-09-06.
Hayward, M.W.; Henschel, P.; O'Brien, J.; Hofmeyr, M.; Balme, G. & Kerley, G. I. H. (2006). "Prey preferences of the leopard (Panthera pardus)" (PDF). Journal of Zoology. 270 (4): 298–313. doi:10.1111/j.1469-7998.2006.00139.x. Archived (PDF) from the original on 2012-11-05.
Schaller, G. (1972). Serengeti: a kingdom of predators. New York: Knopf. ISBN 978-0-394-47242-3. Archived from the original on 2024-05-05. Retrieved 2020-09-18.
"Leopard (Panthera pardus); Physical characteristics and distribution". Comparative Mammalian Brain Collections. Archived from the original on 2011-01-03. Retrieved 2008-06-07.
Salisbury, S. (2014). "Roxanne, oldest spotted leopard in captivity, dies at Acreage preserve". The Palm Beach Post. Archived from the original on 2014-08-11.
Rozhnov, V.V.; Yachmennikova, A.A.; Dronova, N.A.; Naidenko, S.V.; Hernandez-Blanco, J.A.; Chistopolova, M.D.; Pkhitikov, A.B.; Tembotova, F.A.; Trepet, S.A. & Chestin, I.E. (2022). "Experience of the leopard re-covering through reintroduction in the Russian Caucasus" (PDF). Cat News (Special issue): 67–71. Archived (PDF) from the original on 2023-11-11. Retrieved 2023-11-11.
Raza, R.H.; Chauhan, D.S.; Pasha, M.K.S. & Sinha, S. (2012). Illuminating the blind spot: A study on illegal trade in Leopard parts in India (2001–2010) (PDF) (Report). New Delhi: TRAFFIC India, WWF India. Archived (PDF) from the original on 2020-09-24.
Bergin, D. & Nijman, V. (2014). "Open, Unregulated Trade in Wildlife in Morocco's Markets". TRAFFIC Bulletin. 26 (1): 65–70. Archived from the original on 2018-10-31. Retrieved 2015-03-23.
Bergin, D. & Nijman, V. (2015). "Potential benefits of impending Moroccan wildlife trade laws, a case study in carnivore skins". Biodiversity and Conservation. 25 (1): 199–201. doi:10.1007/s10531-015-1042-1. S2CID 34533018. Archived from the original on 2020-01-07. Retrieved 2017-01-11.
Äbischer, T.; Ibrahim, T.; Hickisch, R.; Furrer, R. D.; Leuenberger, C. & Wegmann, D. (2020). "Apex predators decline after an influx of pastoralists in former Central African Republic hunting zones" (PDF). Biological Conservation. 241 108326. Bibcode:2020BCons.24108326A. doi:10.1016/j.biocon.2019.108326. S2CID 213766740. Archived (PDF) from the original on 2020-10-03.
Murphey, R. (1951). "The Decline of North Africa Since the Roman Occupation: Climatic or Human?" (PDF). Annals of the Association of American Geographers. XLI (2): 116–132. doi:10.1080/00045605109352048. Archived (PDF) from the original on 2006-09-14.
"Benin: an African kingdom" (PDF). London: British Museum. Archived (PDF) from the original on 2008-08-05. Retrieved 2016-03-29.
Kipling, R. (1902). "How the Leopard Got His Spots" (PDF). Just So Stories. Macmillan. Archived (PDF) from the original on 2023-11-29. Retrieved 2023-05-14.
Haist, M. (1999). "The Lion, bloodline, and kingship". In Hassig, D. (ed.). The Mark of the Beast: The Medieval Bestiary in Art, Life, and Literature. London: Taylor & Francis. pp. 3–16. ISBN 978-0-8153-2952-7. Archived from the original on 2024-05-05. Retrieved 2021-09-07.
Owen, J. (2005). "Medieval Lion Skulls Reveal Secrets of Tower of London 'Zoo'". National Geographic Magazine. Archived from the original on November 5, 2005. Retrieved 2007-09-05.

wisc.edu (Global: 891^st place; English: 533^rd place)

faculty.nelson.wisc.edu

Johnson, A.; Vongkhamheng, C.; Hedemark, M. & Saithongdam, T. (2006). "Effects of human–carnivore conflict on tiger (Panthera tigris) and prey populations in Lao PDR" (PDF). Animal Conservation. 9 (4): 421–430. Bibcode:2006AnCon...9..421J. doi:10.1111/j.1469-1795.2006.00049.x. S2CID 73637721. Archived (PDF) from the original on 2017-08-10.

worldcat.org (Global: 4^th place; English: 4^th place)

search.worldcat.org

Wozencraft, W. C. (2005). "Species Panthera pardus". In Wilson, D. E.; Reeder, D. M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Johns Hopkins University Press. p. 547. ISBN 978-0-8018-8221-0. OCLC 62265494.
Sauqué, Víctor; Rabal-Garcés, Raquel; Sola-Almagro, Cristina; Cuenca-Bescós, Gloria (18 March 2014). Rook, Lorenzo (ed.). "Bone Accumulation by Leopards in the Late Pleistocene in the Moncayo Massif (Zaragoza, NE Spain)". PLoS ONE. 9 (3) e92144. Bibcode:2014PLoSO...992144S. doi:10.1371/journal.pone.0092144. ISSN 1932-6203. PMC 3958443. PMID 24642667.
Blanco, R. E.; Luzardo-Vera, D. E. (2021). "A biomechanical model to assess the injury risk of leopards (Panthera pardus) hunting by free falling from trees". Biological Communications. 66 (3). doi:10.21638/spbu03.2021.305. hdl:20.500.12008/38512. ISSN 2587-5779.

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Johnson, W. E.; Eizirik, E.; Pecon-Slattery, J.; Murphy, W. J.; Antunes, A.; Teeling, E. & O'Brien, S. J. (2006). "The late Miocene radiation of modern Felidae: a genetic assessment". Science. 311 (5757): 73–77. Bibcode:2006Sci...311...73J. doi:10.1126/science.1122277. PMID 16400146. S2CID 41672825. Archived from the original on 2020-10-04. Retrieved 2019-07-05.

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