Razafimandimbison, Sylvain G. & Rydin, Catarina (2019). "Molecular-based assessments of tribal and generic limits and relationships in Rubiaceae (Gentianales): Polyphyly of Pomazoteae and paraphyly of Ophiorrhizeae and Ophiorrhiza". Taxon. 68 (1): 72–91. doi:10.1002/tax.12023. S2CID146054081.
Razafimandimbison, Sylvain G. & Rydin, Catarina (2019). "Molecular-based assessments of tribal and generic limits and relationships in Rubiaceae (Gentianales): Polyphyly of Pomazoteae and paraphyly of Ophiorrhizeae and Ophiorrhiza". Taxon. 68 (1): 72–91. doi:10.1002/tax.12023. S2CID146054081.