Smythe, L.; Adler, B.; Hartskeerl, R. A.; Galloway, R. L.; Turenne, C. Y.; Levett, P. N. (14 September 2012). "Classification of Leptospira genomospecies 1, 3, 4 and 5 as Leptospira alstonii sp. nov., Leptospira vanthielii sp. nov., Leptospira terpstrae sp. nov. and Leptospira yanagawae sp. nov., respectively". International Journal of Systematic and Evolutionary Microbiology. 63 (Pt 5): 1859–1862. doi:10.1099/ijs.0.047324-0. PMID22984140.
Smythe, L.; Adler, B.; Hartskeerl, R. A.; Galloway, R. L.; Turenne, C. Y.; Levett, P. N. (14 September 2012). "Classification of Leptospira genomospecies 1, 3, 4 and 5 as Leptospira alstonii sp. nov., Leptospira vanthielii sp. nov., Leptospira terpstrae sp. nov. and Leptospira yanagawae sp. nov., respectively". International Journal of Systematic and Evolutionary Microbiology. 63 (Pt 5): 1859–1862. doi:10.1099/ijs.0.047324-0. PMID22984140.