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Heinze, Jurgen; Foitzik, Susanne; Fischer, Birgit; Wanke, Tina; Kipyatkov, Vladilen E. (2003). "The significance of latitudinal variation in body size in a holarctic ant, Leptothorax acervorum". Ecography. 26 (3): 349–55. doi:10.1034/j.1600-0587.2003.03478.x. JSTOR3683375.
Hammond, R. L.; Bourke, A. F. G.; Bruford, M. W. (2001). "Mating frequency and mating system of the polygynous ant Leptothorax acervorum". Molecular Ecology. 10 (11): 2719–28. doi:10.1046/j.0962-1083.2001.01394.x. PMID11883885. S2CID23298932.
Bourke, Andrew F. G. (1991). "Queen behaviour, reproduction and egg cannibalism in multiple-queen colonies of the ant Leptothorax acervorum". Animal Behaviour. 42 (2): 295–310. doi:10.1016/S0003-3472(05)80561-5. S2CID53168799.
Felke, M.; Buschinger, A. (1999). "Social organization, reproductive behavior and ecology of Leptothorax acervorum (Hymenoptera, Formicidae) from the Sierra de Albarracin in central Spain". Insectes Sociaux. 46: 84–91. doi:10.1007/s000400050117. S2CID39506456.
Hammond, R. L.; Bourke, A. F. G.; Bruford, M. W. (2001). "Mating frequency and mating system of the polygynous ant Leptothorax acervorum". Molecular Ecology. 10 (11): 2719–28. doi:10.1046/j.0962-1083.2001.01394.x. PMID11883885. S2CID23298932.