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Pitra, Christian; Dietmar Lieckfeldt; Sylke Frahnert; Joerns Fickel (2002). "Phylogenetic Relationships and Ancestral Areas of the Bustards (Gruiformes: Otididae), Inferred from Mitochondrial DNA and Nuclear Intron Sequences". Molecular Phylogenetics and Evolution. 23 (1): 63–74. Bibcode:2002MolPE..23...63P. doi:10.1006/mpev.2001.1078. PMID12182403.