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Willows-Munro, S.; Robinson, T. J.; Matthee, C. A. (June 2005). "Utility of nuclear DNA intron markers at lower taxonomic levels: Phylogenetic resolution among nine Tragelaphus spp". Molecular Phylogenetics and Evolution. 35 (3): 624–36. doi:10.1016/j.ympev.2005.01.018. PMID15878131.
Hassanin, A.; Delsuc, F.; Ropiquet, A.; Hammer, C.; Jansen van Vuuren, B.; Matthee, C.; Ruiz-Garcia, M.; Catzeflis, F.; Areskoug, V.; Nguyen, T.T.; Couloux, A. (2012). "Pattern and timing of diversification of Cetartiodactyla (Mammalia, Laurasiatheria), as revealed by a comprehensive analysis of mitochondrial genomes". Comptes Rendus Biologies. 335 (1): 32–50. doi:10.1016/j.crvi.2011.11.002. PMID22226162.
Benirschke, K.; Ruedi, D.; Muller, H.; Kumamoto, A.T.; Wagner, K.L.; Downes, H.S. (1980). "The unusual karyotype of the lesser kudu,Tragelaphus imberbis". Cytogenetic and Genome Research. 26 (2–4): 85–92. doi:10.1159/000131429. PMID7389415.