Artiss T, Schultz TR, Polhemus DA, Simon C (2001). "Molecular phylogenetic analysis of the dragonfly genera Libellula, Ladona, and Plathemis (Odonata: Libellulidae) based on mitochondrial cytochrome oxidase I and 16S rRNA sequence data". Molecular Phylogenetics and Evolution. 18 (3): 348–61. doi:10.1006/mpev.2000.0867. PMID11277629.
Artiss T, Schultz TR, Polhemus DA, Simon C (2001). "Molecular phylogenetic analysis of the dragonfly genera Libellula, Ladona, and Plathemis (Odonata: Libellulidae) based on mitochondrial cytochrome oxidase I and 16S rRNA sequence data". Molecular Phylogenetics and Evolution. 18 (3): 348–61. doi:10.1006/mpev.2000.0867. PMID11277629.