Boulenger, G. A. (1899). "Second contribution to the ichthyology of Lake Tanganyika.-- On the fishes obtained by the Congo Free State Expedition under Lieut. Lemaire in 1898". Transactions of the Zoological Society of London. 15 (4(1)): 87–96. doi:10.1111/j.1096-3642.1899.tb00020.x.