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Bruche, S.; Gusset, M.; Lippold, S.; Barnett, R.; Eulenberger, K.; Junhold, J.; Driscoll, C. A.; Hofreiter, M. (2012). "A genetically distinct lion (Panthera leo) population from Ethiopia". European Journal of Wildlife Research. 59 (2): 215–225. doi:10.1007/s10344-012-0668-5. S2CID508478.
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Dollinger, P.; Geser, S. "Lion: In the Zoo (subpage)". Visit the Zoo. WAZA (World Association of Zoos and Aquariums). Archived from the original on 29 September 2011. Retrieved 5 April 2011.