R.T. Chesser; S.M. Billerman; K.J. Burns; C. Cicero; J.L. Dunn; B.E. Hernández-Baños; R.A. Jiménez; A.W. Kratter; N.A. Mason; P.C. Rasmussen; J.V. Remsen, Jr.; D.F. Stotz; K. Winker (July 29, 2022). "Check-list of North and Middle American Birds". American Ornithological Society. Retrieved July 7, 2022.{{cite web}}: CS1 maint: date and year (link)
doi.org
G.E. Alvarado; A. Bonilla-Mata; G. Cárdenas-Sandí (2024). "A Synopsis of the Geology, Geotectonic, and Geological History of Costa Rica". In A. Quesada-Román (ed.). Landscapes and Landforms of Costa Rica. World Geommorphological Landscapes. Springer, Chamb. pp. 3–33. doi:10.1007/978-3-031-64940-0_1. ISBN978-3-031-64940-0.
J.F. Ornelas; C. González; A. Espinoisa de los Monteros; F. Rodríguez-Gómez; L.M. García-Feria (2014). "In and out of Mesoamerica: Temporal divergence of Amazilia hummingbirds pre-dates the orthodox account of the completion of the Isthmus of Panama". Journal of Biogeography. 41 (1): 168–181. doi:10.1111/jbi.12184.