Nathan K Lujan; Jonathan W Armbruster; Nathan R Lovejoy (December 2018). "Multilocus phylogeny, diagnosis and generic revision of the Guiana Shield endemic suckermouth armoured catfish tribe Lithoxini (Loricariidae: Hypostominae)". Zoological Journal of the Linnean Society. 184 (4): 1169–1186. doi:10.1093/zoolinnean/zly025.