Federle, W.; Maschwitz, U.; Fiala, B. (1998). "The two-partner ant-plant system of Camponotus (Colobopsis) sp. 1 and Macaranga puncticulata (Euphorbiaceae): Natural history of the exceptional ant partner". Insectes Sociaux. 45 (1): 1–16. doi:10.1007/s000400050064. S2CID26704950.
Federle, W.; Maschwitz, U.; Fiala, B. (1998). "The two-partner ant-plant system of Camponotus (Colobopsis) sp. 1 and Macaranga puncticulata (Euphorbiaceae): Natural history of the exceptional ant partner". Insectes Sociaux. 45 (1): 1–16. doi:10.1007/s000400050064. S2CID26704950.