Leneveu, Julien; Chichvarkhin, Anton; Wahlberg, Niklas (1 June 2009). "Varying rates of diversification in the genus Melitaea (Lepidoptera: Nymphalidae) during the past 20 million years". Biological Journal of the Linnean Society. 97 (2): 346–361. doi:10.1111/j.1095-8312.2009.01208.x. ISSN0024-4066.
Tóth, János P.; Varga, Zoltán (2011-09-01). "Inter- and intraspecific variation in the genitalia of the 'Melitaea phoebe group' (Lepidoptera, Nymphalidae)". Zoologischer Anzeiger. 250 (3): 258–268. Bibcode:2011ZooAn.250..258T. doi:10.1016/j.jcz.2011.05.002.
Tóth, János P.; Varga, Katalin; Végvári, Zsolt; Varga, Zoltán (2013-04-01). "Distribution of the eastern knapweed fritillary (Melitaea ornata Christoph, 1893) (Lepidoptera: Nymphalidae): past, present and future". Journal of Insect Conservation. 17 (2): 245–255. doi:10.1007/s10841-012-9503-2. ISSN1366-638X. S2CID254595156.
harvard.edu
ui.adsabs.harvard.edu
Tóth, János P.; Varga, Zoltán (2011-09-01). "Inter- and intraspecific variation in the genitalia of the 'Melitaea phoebe group' (Lepidoptera, Nymphalidae)". Zoologischer Anzeiger. 250 (3): 258–268. Bibcode:2011ZooAn.250..258T. doi:10.1016/j.jcz.2011.05.002.
Tóth, János P.; Varga, Katalin; Végvári, Zsolt; Varga, Zoltán (2013-04-01). "Distribution of the eastern knapweed fritillary (Melitaea ornata Christoph, 1893) (Lepidoptera: Nymphalidae): past, present and future". Journal of Insect Conservation. 17 (2): 245–255. doi:10.1007/s10841-012-9503-2. ISSN1366-638X. S2CID254595156.
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Leneveu, Julien; Chichvarkhin, Anton; Wahlberg, Niklas (1 June 2009). "Varying rates of diversification in the genus Melitaea (Lepidoptera: Nymphalidae) during the past 20 million years". Biological Journal of the Linnean Society. 97 (2): 346–361. doi:10.1111/j.1095-8312.2009.01208.x. ISSN0024-4066.