Lehr, E. (2006). "Taxonomic status of some species of Peruvian Phrynopus (Anura: Leptodactylidae), with the description of a new species from the Andes of southern Peru". Herpetologica. 62 (3): 331–347. doi:10.1655/0018-0831(2006)62[331:tsosso]2.0.co;2. JSTOR3893490.
Lehr, E. (2006). "Taxonomic status of some species of Peruvian Phrynopus (Anura: Leptodactylidae), with the description of a new species from the Andes of southern Peru". Herpetologica. 62 (3): 331–347. doi:10.1655/0018-0831(2006)62[331:tsosso]2.0.co;2. JSTOR3893490.