Manns U, Bremer B (2010). "Towards a better understanding of intertribal relationships and stable tribal delimitations within Cinchonoideae s.s. (Rubiaceae)". Molecular Phylogenetics and Evolution. 56 (1): 21–39. doi:10.1016/j.ympev.2010.04.002. PMID20382247.
Löfstrand SD, Krüger Å, Razafimandimbison SG, Bremer B (2014). "Phylogeny and generic delimitations in the sister tribes Hymenodictyeae and Naucleeae (Rubiaceae)". Systematic Botany. 39: 304–315. doi:10.1600/036364414X678116.
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Manns U, Bremer B (2010). "Towards a better understanding of intertribal relationships and stable tribal delimitations within Cinchonoideae s.s. (Rubiaceae)". Molecular Phylogenetics and Evolution. 56 (1): 21–39. doi:10.1016/j.ympev.2010.04.002. PMID20382247.
