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Catanese, Gaetano; Manchado, Manuel; Infante, Carlos (15 February 2010). "Evolutionary relatedness of mackerels of the genus Scomber based on complete mitochondrial genomes: Strong support to the recognition of Atlantic Scomber colias and Pacific Scomber japonicus as distinct species". Gene. 452 (1): 35–43. doi:10.1016/j.gene.2009.12.004. PMID20035845.