Klaus, M., Moore, R., & Vyse, E. (2001, July). Microgeographic variation in allozymes and mitochondrial DNA of Microtus richardsoni, the water vole, in the Beartooth Mountains of Montana and Wyoming, U.S.A. Canadian Journal of Zoology, 79(7), 1286-1295. doi:10.1139/z01-082
Ludwig, D., & Anderson, P. (2009). Metapopulation Biology: Microtus richardsoni in the Rocky Mountain Front Range of Alberta. Northwestern Naturalist, 90(1), 1-16. Retrieved March 20, 2015, from https://www.jstor.org/stable/20628107