OSWALD, JOHN D. (October 1996). "A new brachypterous Nusalala species from Costa Rica, with comments on the evolution of flightlessness in brown lacewings (Neuroptera: Hemerobiidae)". Systematic Entomology. 21 (4): 343–352. doi:10.1111/j.1365-3113.1996.tb00603.x.