Orlando, L.; Ginolhac, A.; Zhang, G.; Froese, D.; Albrechtsen, A.; Stiller, M.; Schubert, M.; Cappellini, E.; Petersen, B.; et al. (4 July 2013). "Recalibrating Equus evolution using the genome sequence of an early Middle Pleistocene horse". Nature. 499 (7456): 74–8. Bibcode:2013Natur.499...74O. doi:10.1038/nature12323. PMID23803765. S2CID4318227.
Stimpson, C. M.; Lister, A.; Parton, Ash; Clark-Balzan, Laine; Breeze, Paul S.; Drake, Nick A.; Groucutt, H. S.; Jennings, R.; Scerri, E. M.L.; White, T. S.; Zahir, M.; Duval, M.; Grün, R.; Al-Omari, A.; Al Murayyi, K. S. M.; Zalmout, I. S.; Mufarreh, Y. A.; Memesh, A. M.; Petraglia, M. D. (2016). "Middle Pleistocene vertebrate fossils from the Nefud Desert, Saudi Arabia: Implications for biogeography and palaeoecology". Quaternary Science Reviews. 143: 13–36. doi:10.1016/j.quascirev.2016.05.016. hdl:10072/142575.
Stimpson, C. M.; Lister, A.; Parton, Ash; Clark-Balzan, Laine; Breeze, Paul S.; Drake, Nick A.; Groucutt, H. S.; Jennings, R.; Scerri, E. M.L.; White, T. S.; Zahir, M.; Duval, M.; Grün, R.; Al-Omari, A.; Al Murayyi, K. S. M.; Zalmout, I. S.; Mufarreh, Y. A.; Memesh, A. M.; Petraglia, M. D. (2016). "Middle Pleistocene vertebrate fossils from the Nefud Desert, Saudi Arabia: Implications for biogeography and palaeoecology". Quaternary Science Reviews. 143: 13–36. doi:10.1016/j.quascirev.2016.05.016. hdl:10072/142575.
Orlando, L.; Ginolhac, A.; Zhang, G.; Froese, D.; Albrechtsen, A.; Stiller, M.; Schubert, M.; Cappellini, E.; Petersen, B.; et al. (4 July 2013). "Recalibrating Equus evolution using the genome sequence of an early Middle Pleistocene horse". Nature. 499 (7456): 74–8. Bibcode:2013Natur.499...74O. doi:10.1038/nature12323. PMID23803765. S2CID4318227.
Orlando, L.; Ginolhac, A.; Zhang, G.; Froese, D.; Albrechtsen, A.; Stiller, M.; Schubert, M.; Cappellini, E.; Petersen, B.; et al. (4 July 2013). "Recalibrating Equus evolution using the genome sequence of an early Middle Pleistocene horse". Nature. 499 (7456): 74–8. Bibcode:2013Natur.499...74O. doi:10.1038/nature12323. PMID23803765. S2CID4318227.
Orlando, L.; Ginolhac, A.; Zhang, G.; Froese, D.; Albrechtsen, A.; Stiller, M.; Schubert, M.; Cappellini, E.; Petersen, B.; et al. (4 July 2013). "Recalibrating Equus evolution using the genome sequence of an early Middle Pleistocene horse". Nature. 499 (7456): 74–8. Bibcode:2013Natur.499...74O. doi:10.1038/nature12323. PMID23803765. S2CID4318227.