Reina-Rodríguez, Guillermo A.; Bogarín, Diego; Hernandez, Yerlin; Nicholls-Giraldo, Isabel & Pérez-Escobar, Oscar A. (2023), "A new Ophidion (Orchidaceae, Pleurothallidinae) from the Pacific lowlands of Colombia and the unresolved phylogenetic position of Phloeophila s.l.", Systematics and Biodiversity, 21 (1): 2160504, doi:10.1080/14772000.2022.2160504