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Devey, D. S.; Bateman, R.M.; Fay, M.F. & Hawkins, J.A. (2008), "Friends or Relatives? Phylogenetics and Species Delimitation in the Controversial European Orchid Genus Ophrys", Annals of Botany, 101 (3): 385–402, doi:10.1093/aob/mcm299, PMC2701817, PMID18184645
Gögler, J.; Stökl, J.; Sramkova, A.; Twele, R.; Francke, W.; Cozzolino, S.; Cortis, P.; Scrugli, A.; Ayasse, M. (2009). "Ménage à Trois — Two Endemic Species of Deceptive Orchids and One Pollinator Species". Evolution. 63 (9): 2222–2234. doi:10.1111/j.1558-5646.2009.00712.x. PMID19473395.
Devey, D. S.; Bateman, R.M.; Fay, M.F. & Hawkins, J.A. (2008), "Friends or Relatives? Phylogenetics and Species Delimitation in the Controversial European Orchid Genus Ophrys", Annals of Botany, 101 (3): 385–402, doi:10.1093/aob/mcm299, PMC2701817, PMID18184645
ncbi.nlm.nih.gov
Devey, D. S.; Bateman, R.M.; Fay, M.F. & Hawkins, J.A. (2008), "Friends or Relatives? Phylogenetics and Species Delimitation in the Controversial European Orchid Genus Ophrys", Annals of Botany, 101 (3): 385–402, doi:10.1093/aob/mcm299, PMC2701817, PMID18184645