Borg-Karlson, Anna-Karin; Inga Groth; Lennart Ågren & Bertil Kullenberg (1993). "Form-specific fragances from Ophrys insectifera L. (Orchidaceae) attract species of different pollinator genera. Evidence of sympatric speciation?". Chemoecology. 4 (1). Birkhäuser Basel: 39–45. doi:10.1007/BF01245895. S2CID26694626.
Borg-Karlson, Anna-Karin; Inga Groth; Lennart Ågren & Bertil Kullenberg (1993). "Form-specific fragances from Ophrys insectifera L. (Orchidaceae) attract species of different pollinator genera. Evidence of sympatric speciation?". Chemoecology. 4 (1). Birkhäuser Basel: 39–45. doi:10.1007/BF01245895. S2CID26694626.