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Buston P (2004). "Does the presence of non-breeders enhance the fitness of breeders? An experimental analysis in the clown anemonefish Amphiprion percula". Behavioral Ecology and Sociobiology. 57 (1): 23–31. Bibcode:2004BEcoS..57...23B. doi:10.1007/s00265-004-0833-2. S2CID24516887.