Sakai, Shoko; Tamiji, Inoue (1 January 1999). "A new pollination system: dung-beetle pollination discovered in Orchidantha inouei (Lowiaceae, Zingiberales) in Sarawak, Malaysia". American Journal of Botany. 86 (1): 56–61. doi:10.2307/2656954. JSTOR2656954. PMID21680345.
Sakai, Shoko; Tamiji, Inoue (1 January 1999). "A new pollination system: dung-beetle pollination discovered in Orchidantha inouei (Lowiaceae, Zingiberales) in Sarawak, Malaysia". American Journal of Botany. 86 (1): 56–61. doi:10.2307/2656954. JSTOR2656954. PMID21680345.
Stevens, P.F. "Lowiaceae". Angiosperm Phylogeny Website. Retrieved 2013-08-20.
nih.gov
pubmed.ncbi.nlm.nih.gov
Sakai, Shoko; Tamiji, Inoue (1 January 1999). "A new pollination system: dung-beetle pollination discovered in Orchidantha inouei (Lowiaceae, Zingiberales) in Sarawak, Malaysia". American Journal of Botany. 86 (1): 56–61. doi:10.2307/2656954. JSTOR2656954. PMID21680345.