Andreas R.Dunz & Ulrich K.Schliewen (2013). "Molecular phylogeny and revised classification of the haplotilapiine cichlid fishes formerly referred to as Tilapia". Molecular Phylogenetics and Evolution. 68 (1): 64–80. doi:10.1016/j.ympev.2013.03.015. PMID23542002.
Nagl, Sandra; Tichy, Herbert; Mayer, Werner E.; et al. (2001). "Classification and Phylogenetic Relationships of African Tilapiine Fishes Inferred from Mitochondrial DNA Sequences". Molecular Phylogenetics and Evolution. 20 (3): 361–374. doi:10.1006/mpev.2001.0979. PMID11527464.
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Andreas R.Dunz & Ulrich K.Schliewen (2013). "Molecular phylogeny and revised classification of the haplotilapiine cichlid fishes formerly referred to as Tilapia". Molecular Phylogenetics and Evolution. 68 (1): 64–80. doi:10.1016/j.ympev.2013.03.015. PMID23542002.
Nagl, Sandra; Tichy, Herbert; Mayer, Werner E.; et al. (2001). "Classification and Phylogenetic Relationships of African Tilapiine Fishes Inferred from Mitochondrial DNA Sequences". Molecular Phylogenetics and Evolution. 20 (3): 361–374. doi:10.1006/mpev.2001.0979. PMID11527464.