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Cole, Charles J.; Townsend, Carol R.; Reynolds, Robert P.; MacCulloch, Ross D. & Lathrop, Amy (2013). "Amphibians and reptiles of Guyana, South America: illustrated keys, annotated species accounts, and a biogeographic synopsis". Proceedings of the Biological Society of Washington. 125 (4): 317–578. doi:10.2988/0006-324X-125.4.317. S2CID86665287.
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Lahanas, Pete N. & Savage, Jay M. (1992). "A new species of caecilian from the Península de Osa of Costa Rica". Copeia. 1992 (3): 703–708. doi:10.2307/1446146. JSTOR1446146.
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Cole, Charles J.; Townsend, Carol R.; Reynolds, Robert P.; MacCulloch, Ross D. & Lathrop, Amy (2013). "Amphibians and reptiles of Guyana, South America: illustrated keys, annotated species accounts, and a biogeographic synopsis". Proceedings of the Biological Society of Washington. 125 (4): 317–578. doi:10.2988/0006-324X-125.4.317. S2CID86665287.