Anderson, R. P.; Gutiérrez, E. C. E. (2009). "Chapter 2. Taxonomy, Distribution, and Natural History of the Genus Heteromys (Rodentia: Heteromyidae) in Central and Eastern Venezuela, with the Description of a New Species from the Cordillera de la Costa". Bulletin of the American Museum of Natural History. 331: 33–93. doi:10.1206/582-2.1. S2CID41011596.
Anderson, R. P.; Gutiérrez, E. C. E. (2009). "Chapter 2. Taxonomy, Distribution, and Natural History of the Genus Heteromys (Rodentia: Heteromyidae) in Central and Eastern Venezuela, with the Description of a New Species from the Cordillera de la Costa". Bulletin of the American Museum of Natural History. 331: 33–93. doi:10.1206/582-2.1. S2CID41011596.