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Tumenta, P. N.; Kok, J. S.; Van Rijssel, J. C.; Buij, R.; Croes, B. M.; Funston, P. J.; De Iongh, H. H.; Udo de Haes, H. A. (2010). "Threat of rapid extermination of the lion (Panthera leo leo) in Waza National Park, Northern Cameroon". African Journal of Ecology. 48 (4): 888−894. doi:10.1111/j.1365-2028.2009.01181.x. hdl:1887/14372. S2CID56451273.
Dubach, J. M.; Briggs, M. B.; White, P. A.; Ament, B. A.; Patterson, B. D. (2013). "Genetic perspectives on "Lion Conservation Units" in Eastern and Southern Africa". Conservation Genetics. 14 (4): 741−755. doi:10.1007/s10592-013-0453-3. S2CID751286.
Tumenta, P. N.; Kok, J. S.; Van Rijssel, J. C.; Buij, R.; Croes, B. M.; Funston, P. J.; De Iongh, H. H.; Udo de Haes, H. A. (2010). "Threat of rapid extermination of the lion (Panthera leo leo) in Waza National Park, Northern Cameroon". African Journal of Ecology. 48 (4): 888−894. doi:10.1111/j.1365-2028.2009.01181.x. hdl:1887/14372. S2CID56451273.
Wilson, R. T. (1979). "Wildlife in Southern Darfur, Sudan: Distribution and status at present and in the recent past". Mammalia. 43 (3): 323−338. doi:10.1515/mamm.1979.43.3.323. S2CID85228859.
Bauer, H.; de Iongh, H.; Sogbohossou, E. (2010). "Assessment and mitigation of human-lion conflict in West and Central Africa". Mammalia. 74 (4): 363–367. doi:10.1515/MAMM.2010.048. S2CID86228533.
Bruche, S.; Gusset, M.; Lippold, S.; Barnett, R.; Eulenberger, K.; Junhold, J.; Driscoll, C. A.; Hofreiter, M. (2012). "A genetically distinct lion (Panthera leo) population from Ethiopia". European Journal of Wildlife Research. 59 (2): 215–225. doi:10.1007/s10344-012-0668-5. S2CID508478.
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Kitchener, A. C.; Breitenmoser-Würsten, C.; Eizirik, E.; Gentry, A.; Werdelin, L.; Wilting, A.; Yamaguchi, N.; Abramov, A. V.; Christiansen, P.; Driscoll, C.; Duckworth, J. W.; Johnson, W.; Luo, S.-J.; Meijaard, E.; O'Donoghue, P.; Sanderson, J.; Seymour, K.; Bruford, M.; Groves, C.; Hoffmann, M.; Nowell, K.; Timmons, Z. & Tobe, S. (2017). "A revised taxonomy of the Felidae: The final report of the Cat Classification Task Force of the IUCN Cat Specialist Group"(PDF). Cat News (Special Issue 11): 71−73.