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Asad, M.; Martoni, F.; Ross, J.G.; Waseem, M. & Paterson, A.M. (2019). "Assessing subspecies status of leopards (Panthera pardus) of northern Pakistan using mitochondrial DNA". PeerJ. 7: e7243. doi:10.7717/peerj.7243. hdl:10182/10848. PMC 6640621. PMID 31341733.
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harvard.edu

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Miththapala, S.; Seidensticker, J. & O'Brien, S. J. (1996). "Phylogeographic Subspecies Recognition in Leopards (P. pardus): Molecular Genetic Variation". Conservation Biology. 10 (4): 1115–1132. Bibcode:1996ConBi..10.1115M. doi:10.1046/j.1523-1739.1996.10041115.x.
Uphyrkina, O.; Johnson, E.W.; Quigley, H.; Miquelle, D.; Marker, L.; Bush, M. & O'Brien, S. J. (2001). "Phylogenetics, genome diversity and origin of modern leopard, Panthera pardus" (PDF). Molecular Ecology. 10 (11): 2617–2633. Bibcode:2001MolEc..10.2617U. doi:10.1046/j.0962-1083.2001.01350.x. PMID 11883877. S2CID 304770. Archived from the original (PDF) on 28 April 2020. Retrieved 5 March 2021.
Khorozyan, I. (2014). "Morphological variation and sexual dimorphism of the common leopard (Panthera pardus) in the Middle East and their implications for species taxonomy and conservation". Mammalian Biology – Zeitschrift für Säugetierkunde. 79 (6): 398–405. Bibcode:2014MamBi..79..398K. doi:10.1016/j.mambio.2014.07.004.
Miththapala, S.; Seidensticker, J.; O'Brien, S. J. (1996). "Phylogeographic Subspecies Recognition in Leopards (Panthera pardus): Molecular Genetic Variation". Conservation Biology. 10 (4): 1115–1132. Bibcode:1996ConBi..10.1115M. doi:10.1046/j.1523-1739.1996.10041115.x.
Uphyrkina, O.; Johnson, E.W.; Quigley, H.; Miquelle, D.; Marker, L.; Bush, M. & O'Brien, S. J. (2001). "Phylogenetics, genome diversity and origin of modern leopard, Panthera pardus". Molecular Ecology. 10 (11): 2617–2633. Bibcode:2001MolEc..10.2617U. doi:10.1046/j.0962-1083.2001.01350.x. PMID 11883877. S2CID 304770.
Farhadinia, M.S.; Ahmadi, M.; Sharbafi, E.; Khosravi, S.; Alinezhad, H. & Macdonald, D.W. (2015). "Leveraging trans-boundary conservation partnerships: Persistence of Persian leopard (Panthera pardus saxicolor) in the Iranian Caucasus" (PDF). Biological Conservation. 191: 770–778. Bibcode:2015BCons.191..770F. doi:10.1016/j.biocon.2015.08.027.
Farhadinia, M.S.; McClintock, B.T.; Johnson, P.J.; Behnoud, P.; Hobeali, K.; Moghadas, P.; Hunter, L.T. & Macdonald, D.W. (2019). "A paradox of local abundance amidst regional rarity: the value of montane refugia for Persian leopard conservation". Scientific Reports. 9 (1): 14622. Bibcode:2019NatSR...914622F. doi:10.1038/s41598-019-50605-2. PMC 6788991. PMID 31604967.
Farhadinia, M.S.; Johnson, P.J.; Macdonald, D.W. & Hunter, L.T.B. (2018). "Anchoring and adjusting amidst humans: Ranging behavior of Persian leopards along the Iran-Turkmenistan borderland". PLOS ONE. 13 (5): e0196602. Bibcode:2018PLoSO..1396602F. doi:10.1371/journal.pone.0196602. PMC 5931651. PMID 29719005.
Sekercioglu, C.; Anderson, S.; Akcay, E.; Bilgin, R.; Can, O. & Semiz, G. (2011). "Turkey's Globally Important Biodiversity In Crisis" (PDF). Biological Conservation. 144 (12): 2752–2769. Bibcode:2011BCons.144.2752S. doi:10.1016/j.biocon.2011.06.025. S2CID 18094317.
Naderi, M.; Farashi, A. & Erdi, M.A. (2018). "Persian leopard's (Panthera pardus saxicolor) unnatural mortality factors analysis in Iran". PLOS ONE. 13 (4): e0195387. Bibcode:2018PLoSO..1395387N. doi:10.1371/journal.pone.0195387. PMC 5918793. PMID 29694391.
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