A. A. Echelle; P. J. Connor (1989). "Rapid, geographically extensive genetic introgression after secondary contact between two pupfish species (Cyprinodon, Cyprinodontidae)". Evolution. 43 (4): 717–727. doi:10.1111/j.1558-5646.1989.tb05171.x. PMID28564193.
J. A. Rosenfield; S. Nolasco; C. Sandoval; A. Kodric-Brown (2004). "The Role of Hybrid Vigor in the Replacement of Pecos Pupfish by Its Hybrids with Sheepshead Minnow". Conservation Biology. 18 (6): 1589–1598. doi:10.1111/j.1523-1739.2004.00356.x. S2CID39553092.
A. A. Echelle; C. W. Hoagstrom; A. F. Echelle; E. James (1997). "Expanded occurrence of genetically introgressed pupfish ( Cyprinodontidae : Cyprinodon pecosensis X variegatus ) in New Mexico". Southwestern Naturalist. 42 (3): 336–339. JSTOR30055287.
A. A. Echelle; P. J. Connor (1989). "Rapid, geographically extensive genetic introgression after secondary contact between two pupfish species (Cyprinodon, Cyprinodontidae)". Evolution. 43 (4): 717–727. doi:10.1111/j.1558-5646.1989.tb05171.x. PMID28564193.
J. A. Rosenfield; S. Nolasco; C. Sandoval; A. Kodric-Brown (2004). "The Role of Hybrid Vigor in the Replacement of Pecos Pupfish by Its Hybrids with Sheepshead Minnow". Conservation Biology. 18 (6): 1589–1598. doi:10.1111/j.1523-1739.2004.00356.x. S2CID39553092.