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Brehm, Gunnar; Pitkin, Linda M.; Hilt, Nadine; Fiedler, Konrad (2005). "Montane Andean rain forests are a global diversity hotspot of geometrid moths". Journal of Biogeography. 32 (9): 1621–1627. Bibcode:2005JBiog..32.1621B. doi:10.1111/j.1365-2699.2005.01304.x.
Liede-Schumann, S.; Breckle, Siegmar-W. (2008). Provisional checklists of flora and fauna of the San Francisco Valley and its surroundings: Reserva Biológica San Francisco, Prov. Zamora-Chinchipe, Southern Ecuador. Bonn, Germany: Society of Tropical Ecology. ISBN9783980778039. OCLC228007388.
