Guayasamin, Juan M.; Almeida-Reinoso, Diego & Nogales-Sornosa, Fernando (2004). "Two new species of frogs (Leptodactylidae: Eleutherodactylus) from the high Andes of northern Ecuador". Herpetological Monographs. 18: 127–141. doi:10.1655/0733-1347(2004)018[0127:TNSOFL]2.0.CO;2. JSTOR4093022.
Guayasamin, Juan M.; Almeida-Reinoso, Diego & Nogales-Sornosa, Fernando (2004). "Two new species of frogs (Leptodactylidae: Eleutherodactylus) from the high Andes of northern Ecuador". Herpetological Monographs. 18: 127–141. doi:10.1655/0733-1347(2004)018[0127:TNSOFL]2.0.CO;2. JSTOR4093022.