Frost, Darrel R. (2022). "Pristimantis w-nigrum (Boettger, 1892)". Amphibian Species of the World: An Online Reference. Version 6.1. American Museum of Natural History. Retrieved 4 November 2022.
Kieswetter, C. M.; Schneider, C. J. (2013). "Phylogeography in the northern Andes: Complex history and cryptic diversity in a cloud forest frog, Pristimantis w-nigrum (Craugastoridae)". Molecular Phylogenetics and Evolution. 69 (3): 417–429. Bibcode:2013MolPE..69..417K. doi:10.1016/j.ympev.2013.08.007. PMID23978627.
Kieswetter, C. M.; Schneider, C. J. (2013). "Phylogeography in the northern Andes: Complex history and cryptic diversity in a cloud forest frog, Pristimantis w-nigrum (Craugastoridae)". Molecular Phylogenetics and Evolution. 69 (3): 417–429. Bibcode:2013MolPE..69..417K. doi:10.1016/j.ympev.2013.08.007. PMID23978627.
Kieswetter, C. M.; Schneider, C. J. (2013). "Phylogeography in the northern Andes: Complex history and cryptic diversity in a cloud forest frog, Pristimantis w-nigrum (Craugastoridae)". Molecular Phylogenetics and Evolution. 69 (3): 417–429. Bibcode:2013MolPE..69..417K. doi:10.1016/j.ympev.2013.08.007. PMID23978627.