Cherin, Marco; Iurino, Dawid A.; Sardella, Raffaele (2013). "Earliest occurrence of Puma pardoides (Owen, 1846) (Carnivora, Felidae) at the Plio/Pleistocene transition in western Europe: New evidence from the Middle Villafranchian assemblage of Montopoli, Italy". Comptes Rendus Palevol. 12 (3): 165–171. Bibcode:2013CRPal..12..165C. doi:10.1016/j.crpv.2013.01.002.
Hemmer, H.; Kahlike, R. D.; Vekua, A. K. (2004). "The Old World puma Puma pardoides (Owen, 1846) (Carnivora: Felidae) in the Lower Villafranchian (Upper Pliocene) of Kvabebi (East Georgia, Transcaucasia) and its evolutionary and biogeographical significance". Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen. 233: 197–233. doi:10.1127/njgpa/233/2004/197.
Cherin, Marco; Iurino, Dawid A.; Sardella, Raffaele (2013). "Earliest occurrence of Puma pardoides (Owen, 1846) (Carnivora, Felidae) at the Plio/Pleistocene transition in western Europe: New evidence from the Middle Villafranchian assemblage of Montopoli, Italy". Comptes Rendus Palevol. 12 (3): 165–171. Bibcode:2013CRPal..12..165C. doi:10.1016/j.crpv.2013.01.002.