Evans, N.J. (1986). "An investigation of the status of the terrestrial slugs Arion ater ater (L.) and Arion ater rufus (L.) (Mollusca, Gastropoda, Pulmonata) in Britain". Zoologica Scripta. 15 (4): 313–322. doi:10.1111/j.1463-6409.1986.tb00232.x. S2CID84998531.
Hatteland, B.A.; Solhøy, T.; Schander, C.; Skage, M.; Proschwitz, T. von; Noble, L.R. (2015). "Introgression and differentiation of the invasive slug Arion vulgaris from native A. ater". Malacologia. 58 (1–2): 303–321. doi:10.4002/040.058.0210. S2CID84108976.
Soroka, M.; Kozłowski, J.; Wiktor, A.; Kałuski, T. (2009). "Distribution and genetic diversity of the terrestrial slugs Arion lusitanicus Mabille, 1868 and Arion rufus (Linnaeus, 1758) in Poland based on mitochondrial DNA". Folia Biologica (Kraków). 57 (1–2): 71–81. doi:10.3409/fb57_1-2.71-81. PMID19459464.
Allgaier, C. (2015). "How can two soft bodied animals be precisely connected? A miniature quick-connect system in the slugs, Arion lusitanicus and Arion rufus". Journal of Morphology. 276 (6): 631–648. doi:10.1002/jmor.20361. PMID25532726. S2CID29079897.
Soroka, M.; Kozłowski, J.; Wiktor, A.; Kałuski, T. (2009). "Distribution and genetic diversity of the terrestrial slugs Arion lusitanicus Mabille, 1868 and Arion rufus (Linnaeus, 1758) in Poland based on mitochondrial DNA". Folia Biologica (Kraków). 57 (1–2): 71–81. doi:10.3409/fb57_1-2.71-81. PMID19459464.
Allgaier, C. (2015). "How can two soft bodied animals be precisely connected? A miniature quick-connect system in the slugs, Arion lusitanicus and Arion rufus". Journal of Morphology. 276 (6): 631–648. doi:10.1002/jmor.20361. PMID25532726. S2CID29079897.
Rosetta, R. European red slug. Pacific Northwest Nursery IPM. Oregon State University.
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Evans, N.J. (1986). "An investigation of the status of the terrestrial slugs Arion ater ater (L.) and Arion ater rufus (L.) (Mollusca, Gastropoda, Pulmonata) in Britain". Zoologica Scripta. 15 (4): 313–322. doi:10.1111/j.1463-6409.1986.tb00232.x. S2CID84998531.
Hatteland, B.A.; Solhøy, T.; Schander, C.; Skage, M.; Proschwitz, T. von; Noble, L.R. (2015). "Introgression and differentiation of the invasive slug Arion vulgaris from native A. ater". Malacologia. 58 (1–2): 303–321. doi:10.4002/040.058.0210. S2CID84108976.
Allgaier, C. (2015). "How can two soft bodied animals be precisely connected? A miniature quick-connect system in the slugs, Arion lusitanicus and Arion rufus". Journal of Morphology. 276 (6): 631–648. doi:10.1002/jmor.20361. PMID25532726. S2CID29079897.