Dunz, Andreas R.; Schliewen, Ulrich K. (July 2013). "Molecular phylogeny and revised classification of the Haplotilapiine Cichlid fishes formerly referred to as "Tilapia"". Molecular Phylogenetics and Evolution. 68 (1): 64–80. doi:10.1016/j.ympev.2013.03.015. PMID23542002.
Dunz, Andreas R.; Schliewen, Ulrich K. (July 2013). "Molecular phylogeny and revised classification of the Haplotilapiine Cichlid fishes formerly referred to as "Tilapia"". Molecular Phylogenetics and Evolution. 68 (1): 64–80. doi:10.1016/j.ympev.2013.03.015. PMID23542002.
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Poissons d'eaux douces et saumâtres de basse Guinée, ouest de l'Afrique centrale = The fresh and brackish water fishes of Lower Guinea, West-Central Africa. Stiassny, Melanie L. J., Teugels, Guy G., Hopkins, Carl D., American Museum of Natural History. Paris, France: Institut de recherche pour le développement. 2007. ISBN9782709916202. OCLC216917073.{{cite book}}: CS1 maint: others (link)