Cole, Charles J.; Townsend, Carol R.; Reynolds, Robert P.; MacCulloch, Ross D.; Lathrop, Amy (January 2013). "Amphibians and reptiles of Guyana, South America: illustrated keys, annotated species accounts, and a biogeographic synopsis". Proceedings of the Biological Society of Washington. 125 (4): 317–578. doi:10.2988/0006-324X-125.4.317. S2CID86665287.
Cole, Charles J.; Townsend, Carol R.; Reynolds, Robert P.; MacCulloch, Ross D.; Lathrop, Amy (January 2013). "Amphibians and reptiles of Guyana, South America: illustrated keys, annotated species accounts, and a biogeographic synopsis". Proceedings of the Biological Society of Washington. 125 (4): 317–578. doi:10.2988/0006-324X-125.4.317. S2CID86665287.