Pepper, M.; Doughty, P.; Hutchinson, M.N.; Scott Keogh, J. (December 2011). "Ancient drainages divide cryptic species in Australia's arid zone: Morphological and multi-gene evidence for four new species of Beaked Geckos (Rhynchoedura)". Molecular Phylogenetics and Evolution. 61 (3): 810–822. doi:10.1016/j.ympev.2011.08.012. PMID21884806.
Pepper, M.; Doughty, P.; Hutchinson, M.N.; Scott Keogh, J. (December 2011). "Ancient drainages divide cryptic species in Australia's arid zone: Morphological and multi-gene evidence for four new species of Beaked Geckos (Rhynchoedura)". Molecular Phylogenetics and Evolution. 61 (3): 810–822. doi:10.1016/j.ympev.2011.08.012. PMID21884806.