Dana L. Campbell and Naomi E. Pierce (2003): Phylogenetic relationships of the Riodinidae: Implications for the evolution of ant association. In: C. Boggs, P. Ehrlich, W.B. Watt (editors). Butterflies as Model Systems. Chicago University Press: 395-408. downloadArchived 2013-12-03 at the Wayback Machine
Dana L. Campbell and Naomi E. Pierce (2003): Phylogenetic relationships of the Riodinidae: Implications for the evolution of ant association. In: C. Boggs, P. Ehrlich, W.B. Watt (editors). Butterflies as Model Systems. Chicago University Press: 395-408. downloadArchived 2013-12-03 at the Wayback Machine