Pereira, M.E.; Seeligson, M.L.; Macedonia, J.M. (1988). "The behavioral repertoire of the black-and-white ruffed lemur, Varecia variegata variegata (Primates: Lemuridae)". Folia Primatologica. 51 (1): 1–32. doi:10.1159/000156353. PMID3251818.
Macedonia, Joseph M.; Taylor, Linda L.; Macedonia, J.M. (1985). "Subspecific Divergence in a Loud Call of the Ruffed Lemur (Varecia variegata)". American Journal of Primatology. 9 (4): 295–304. doi:10.1002/ajp.1350090406. PMID31979505. S2CID83954920.
Pereira, M.E.; Seeligson, M.L.; Macedonia, J.M. (1988). "The behavioral repertoire of the black-and-white ruffed lemur, Varecia variegata variegata (Primates: Lemuridae)". Folia Primatologica. 51 (1): 1–32. doi:10.1159/000156353. PMID3251818.
Macedonia, Joseph M.; Taylor, Linda L.; Macedonia, J.M. (1985). "Subspecific Divergence in a Loud Call of the Ruffed Lemur (Varecia variegata)". American Journal of Primatology. 9 (4): 295–304. doi:10.1002/ajp.1350090406. PMID31979505. S2CID83954920.
Macedonia, Joseph M.; Taylor, Linda L.; Macedonia, J.M. (1985). "Subspecific Divergence in a Loud Call of the Ruffed Lemur (Varecia variegata)". American Journal of Primatology. 9 (4): 295–304. doi:10.1002/ajp.1350090406. PMID31979505. S2CID83954920.