Osinov, Alexander G.; Volkov, Alexander A.; Mugue, Nikolai S. (2021). "Charrs of the genus Salvelinus (Salmonidae): hybridization, phylogeny and evolution". Hydrobiologia. 848 (3): 705–726. bioRxiv10.1101/817775. doi:10.1007/s10750-020-04478-6. ISSN0018-8158.
Osinov, Alexander G.; Volkov, Alexander A.; Mugue, Nikolai S. (2021). "Charrs of the genus Salvelinus (Salmonidae): hybridization, phylogeny and evolution". Hydrobiologia. 848 (3): 705–726. bioRxiv10.1101/817775. doi:10.1007/s10750-020-04478-6. ISSN0018-8158.