Meik, Jesse M.; Smith, Eric N.; Canseco-Márquez, Luis; Campbell, Jonathan A. (1 January 2006). "New species of the Plectrohyla bistincta group (Hylidae: Hylinae: Hylini) from Oaxaca, Mexico". Journal of Herpetology. 40 (3): 304–309. doi:10.1670/0022-1511(2006)40[304:nsotpb]2.0.co;2. JSTOR4092998.
Meik, Jesse M.; Smith, Eric N.; Canseco-Márquez, Luis; Campbell, Jonathan A. (1 January 2006). "New species of the Plectrohyla bistincta group (Hylidae: Hylinae: Hylini) from Oaxaca, Mexico". Journal of Herpetology. 40 (3): 304–309. doi:10.1670/0022-1511(2006)40[304:nsotpb]2.0.co;2. JSTOR4092998.